A15.Inglish BCEnc. Blauwe Kaas Encyclopedie, Duaal Hermeneuties Kollegium.
Inglish Site.15.
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TO THE THRISE HO-
NOVRABLE AND EVER LY-
VING VERTVES OF SYR PHILLIP
SYDNEY KNIGHT, SYR JAMES JESUS SINGLETON, SYR CANARIS, SYR LAVRENTI BERIA ; AND TO THE
RIGHT HONORABLE AND OTHERS WHAT-
SOEVER, WHO LIVING LOVED THEM,
AND BEING DEAD GIVE THEM
THEIRE DVE.
***
In the beginning there is darkness. The screen erupts in blue, then a cascade of thick, white hexadecimal numbers and cracked language, ?UnusedStk? and ?AllocMem.? Black screen cedes to blue to white and a pair of scales appear, crossed by a sword, both images drawn in the jagged, bitmapped graphics of Windows 1.0-era clip-art?light grey and yellow on a background of light cyan. Blue text proclaims, ?God on tap!?
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Introduction.
Yes i am getting a little Mobi-Literate(ML) by experimenting literary on my Mobile Phone. Peoplecall it Typographical Laziness(TL).
The first accidental entries for the this part of this encyclopedia.
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This is TempleOS V2.17, the welcome screen explains, a ?Public Domain Operating System? produced by Trivial Solutions of Las Vegas, Nevada. It greets the user with a riot of 16-color, scrolling, blinking text; depending on your frame of reference, it might recall ?DESQview, the ?Commodore 64, or a host of early DOS-based graphical user interfaces. In style if not in specifics, it evokes a particular era, a time when the then-new concept of ?personal computing? necessarily meant programming and tinkering and breaking things.
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Index.
67.Synagogue
68.Ants.
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67.Synagogue:Maat, or divine order.
A synagogue, also spelled synagog (from Greek ????????, transliterated synagog?, meaning "assembly"; ??? ???? beth knesset, meaning "house of assembly"; ??? ????? beth t'fila, meaning "house of prayer"; ??? shul; ?????? esnoga; ??? kahal), is a Jewish house of prayer.
Synagogues have a large hall for prayer (the main sanctuary), and may also have smaller rooms for study and sometimes a social hall and offices. Some have a separate room for Torah study, called the beth midrash (Sefaradi/sefardim) "beis midrash (Ashkenazi/Ashkenazim)???? ???? ("House of Study").
Synagogues are consecrated spaces that can be used only for the purpose of prayer; however a synagogue is not necessary for worship. Communal Jewish worship can be carried out wherever ten Jews (a minyan) assemble. Worship can also be carried out alone or with fewer than ten people assembled together. However there are certain prayers that are communal prayers and therefore can be recited only by a minyan. The synagogue does not replace the long-since destroyed Temple in Jerusalem.
Israelis use the Hebrew term beyt knesset (assembly house). Jews of Ashkenazi descent have traditionally used the Yiddish term "shul" (cognate with the German Schule, school) in everyday speech. Spanish and Portuguese Jews call the synagogue an esnoga. Persian Jews and Karaite Jews use the term Kenesa, which is derived from Aramaic, and some Arabic-speaking Jews use knis. Some Reform and Conservative Jews use the word "temple". The Greek word "Synagogue" is a good all-around term, used in English (and German and French), to cover the preceding possibilities.
Although synagogues existed a long time before the destruction of the 2nd Temple in 70 CE, communal worship in the time while the Temple still stood centered around the korbanot ("sacrificial offerings") brought by the kohanim ("priests") in the Holy Temple. The all-day Yom Kippur service, in fact, was an event in which the congregation both observed the movements of the kohen gadol ("the high priest") as he offered the day's sacrifices and prayed for his success.
During the Babylonian captivity (586?537 BCE) the Men of the Great Assembly formalized and standardized the language of the Jewish prayers. Prior to that people prayed as they saw fit, with each individual praying in his or her own way, and there were no standard prayers that were recited. Rabbi Yohanan ben Zakkai, one of the leaders at the end of the Second Temple era, promulgated the idea of creating individual houses of worship in whatever locale Jews found themselves. This contributed to the continuity of the Jewish people by maintaining a unique identity and a portable way of worship despite the destruction of the Temple, according to many historians.
Synagogues in the sense of purpose-built spaces for worship, or rooms originally constructed for some other purpose but reserved for formal, communal prayer, however, existed long before the destruction of the Second Temple. The earliest archaeological evidence for the existence of very early synagogues comes from Egypt, where stone synagogue dedication inscriptions dating from the 3rd century BCE prove that synagogues existed by that date. A synagogue dating from between 75 and 50 BCE has been uncovered at a Hasmonean-era winter palace near Jericho. More than a dozen Second Temple era synagogues have been identified by archaeologists.
Any Jew or group of Jews can build a synagogue. Synagogues have been constructed by ancient Jewish kings, by wealthy patrons, as part of a wide range of human institutions including secular educational institutions, governments, and hotels, by the entire community of Jews living in a particular place, or by sub-groups of Jews arrayed according to occupation, ethnicity (i.e. the Sephardic, Polish or Persian Jews of a town), style of religious observance (i.e., a Reform or an Orthodox synagogue), or by the followers of a particular rabbi.
There is no set blueprint for synagogues and the architectural shapes and interior designs of synagogues vary greatly. In fact, the influence from other local religious buildings can often be seen in synagogue arches, domes and towers.
Historically, synagogues were built in the prevailing architectural style of their time and place. Thus, the synagogue in Kaifeng, China looked very like Chinese temples of that region and era, with its outer wall and open garden in which several buildings were arranged. The styles of the earliest synagogues resembled the temples of other sects of the eastern Roman Empire. The surviving synagogues of medieval Spain are embellished with mudéjar plasterwork. The surviving medieval synagogues in Budapest and Prague are typical Gothic structures.
The emancipation of Jews in European countries not only enabled Jews to enter fields of enterprise from which they were formerly barred, but gave them the right to build synagogues without needing special permissions, synagogue architecture blossomed. Large Jewish communities wished to show not only their wealth but also their newly acquired status as citizens by constructing magnificent synagogues. These were built across Europe and in the United States in all of the historicist or revival styles then in fashion. Thus there were Neoclassical, Neo-Byzantine, Romanesque Revival, Moorish Revival, Gothic Revival, and Greek Revival. There are Egyptian Revival synagogues and even one Mayan Revival synagogue. In the 19th century and early 20th century heyday of historicist architecture, however, most historicist synagogues, even the most magnificent ones, did not attempt a pure style, or even any particular style, and are best described as eclectic.
In the post-war era, synagogue architecture abandoned historicist styles for modernism.
All synagogues contain a bimah, a table from which the Torah is read, and a desk for the prayer leader;
The Torah ark, (Hebrew: Aron Kodesh????? ????) (called the heikhal????? [temple] by Sephardim) is a cabinet in which the Torah scrolls are kept.
The ark in a synagogue is almost always positioned in such a way such that those who face it are facing towards Jerusalem. Thus, sanctuary seating plans in the Western world generally face east, while those east of Israel face west. Sanctuaries in Israel face towards Jerusalem. Occasionally synagogues face other directions for structural reasons; in such cases, some individuals might turn to face Jerusalem when standing for prayers, but the congregation as a whole does not.
The ark is reminiscent of the Ark of the Covenant which held the tablets inscribed with the Ten Commandments. This is the holiest spot in a synagogue, equivalent to the Holy of Holies. The ark is often closed with an ornate curtain, the parochet ?????, which hangs outside or inside the ark doors.
A large, raised, reader's platform called the bimah (????) by Ashkenazim and tebah by Sephardim, where the Torah scroll is placed to be read is a feature of all synagogues. In Sephardi synagogues it is also used as the prayer leader's reading desk.
Other traditional features include a continually lit lamp or lantern, usually electric in contemporary synagogues, called the ner tamid (?? ????), the "Eternal Light", used as a reminder of the western lamp of the menorah of the Temple in Jerusalem, which remained miraculously lit perpetually. Many have an elaborate chair named for the prophet Elijah which is only sat upon during the ceremony of Brit milah. Many synagogues have a large seven-branched candelabrum commemorating the full Menorah. Most contemporary synagogues also feature a lectern for the rabbi.
A synagogue may be decorated with artwork, but in the Rabbinic and Orthodox tradition, three-dimensional sculptures and depictions of the human body are not allowed as these are considered akin to idolatry.
Until the 19th century, an Ashkenazi synagogue, all seats most often faced the 'Torah Ark. In a Sephardi synagogue, seats were usually arranged around the perimeter of the sanctuary, but when the worshipers stood up to pray, everyone faced the Ark. In Ashkenazi synagogues The Torah was read on a reader's table located in the center of the room, while the leader of the prayer service, the Hazzan, stood at his own lectern or table, facing the Ark. In Sephardic synagogues, the table for reading the Torah was commonly placed at the opposite side of the room from the Torah Ark, leaving the center of the floor empty for the use of a ceremonial procession carrying the Torah between the Ark and the reading table.
Denominational differences.
New York's Reform Temple Emanu-El
Orthodox synagogues feature a partition (mechitzah) dividing the men's and women's seating areas, or a separate women's section located on a balcony.
The German Reform movement which arose in the early 19th century made many changes to the traditional look of the synagogue, keeping with its desire to simultaneously stay Jewish yet be accepted by the host culture.
The first Reform synagogue, which opened in Hamburg in 1811, introduced changes that made the synagogue look more like a church. These included: the installation of an organ to accompany the prayers (even on Shabbat, when musical instruments are proscribed by halakha, a choir to accompany the Hazzan, and vestments for the synagogue rabbi to wear.
In following decades, the central reader's table, the Bimah, was moved to the front of the Reform sanctuary?previously unheard-of[citation needed] in Orthodox synagogues. The rabbi now delivered his sermon from the front, much as the Christian ministers delivered their sermons in a church. The synagogue was renamed a "temple", to emphasize that the movement no longer looked forward to the rebuilding of the Temple in Jerusalem.
Synagogue architecture often follows styles in vogue at the place and time of construction. There is no set blueprint for synagogues and the architectural shapes and interior designs of synagogues vary greatly. According to tradition, the Divine Presence (Shekhinah) can be found wherever there is a minyan, a quorum, of ten. A synagogue always contains an ark, called aron ha-kodesh by Ashkenazim and hekhal by Sephardim, where the Torah scrolls are kept.
The ark may be more or less elaborate, even a cabinet not structurally integral to the building or a portable arrangement whereby a Torah is brought into a space temporarily used for worship. There must also be a table from which the Torah is read. The table, called bimah by eastern Ashkenazim, almemmar (or balemmer) by Central and Western Ashkenazim and tebah by Sephardim, where the Torah is read (and from where the services are conducted in Sephardi synagogues) can range from an elaborate platform integral to the building (many early modern synagogues of central Europe featured bimahs with pillars that rose to support the ceiling), to elaborate free-standing raised platforms, to simple tables. A ner tamid, a constantly lit light as a reminder of the constantly lit menorah of the Temple in Jerusalem. Many synagogues, mainly in Ashkenazi communities, feature a pulpit facing the congregation from which to address the assembled. All synagogues require an amud (Hebrew for "post" or "column"), a desk facing the Ark from which the Hazzan (reader, or prayer leader) leads the prayers.
A synagogue may or may not have artwork, synagogues range from simple, unadorned prayer rooms to elaborately decorated buildings in every architectural style.
The synagogue, or if it is a multi-purpose building, prayer sanctuaries within the synagogue, should face towards Jerusalem. Thus sanctuaries in the Western world generally face east, while those east of Israel face west. Sanctuaries in Israel face towards Jerusalem. But this orientation need not be exact, and occasionally synagogues face other directions for structural reasons, in which case the community may face Jerusalem when standing for prayers.
There are only a few Talmudic instructions on how synagogues could be built, namely that they had to have windows and that they had to be taller than other buildings in town, synagogue plans were generally unrestricted, and even these few rules were often disregarded (especially as Jews were often not allowed to build tall buildings). There is no set blueprint for synagogues and the architectural shapes as well as interior designs of synagogues vary greatly.
Touro Synagogue, the oldest synagogue building still standing in the United States, built in Georgian style.
Baroque style W?odawa Synagogue.
Gothic interior of 13th-century Old New Synagogue of Prague.
Historically, synagogues were built in the prevailing architectural style of their time and place. Thus, the synagogue in Kaifeng, China looked very like Chinese temples of that region and era, with its outer wall and open garden in which several buildings were arranged.
Synagogue of the Kaifeng Jewish community in China.
The styles of the earliest synagogues resembled the temples of other sects of the eastern Roman Empire. The synagogues of Morocco are embellished with the colored tilework characteristic of Moroccan architecture. The surviving medieval synagogues in Budapest, Prague and the German lands are typical Gothic structures.
For much of history, the constraints of anti-semitism and the laws of host countries restricting the building of synagogues visible from the street, or forbidding their construction altogether, meant that synagogues were often built within existing buildings, or opening form interior courtyards. In both Europe and in the Muslim world, old synagogues with elaborate interior architecture can be found hidden within nondescript buildings.
Where the building of synagogues was permitted, they were built in the prevailing architectural style of the time and place. Many European cities had elaborate Renaissance synagogues, of which a few survive. In Italy there were many synagogues in the style of the Italian Renaissance (see Leghorn; Padua; and Venice). Those in Padua and Venice possess interiors of great beauty, and are excellent examples of Renaissance work. With the coming of the Baroque era, Baroque synagogues appeared across Europe.
The emancipation of Jews in European countries and of Jews in Muslim countries colonized by European countries gave Jews the right to build large, elaborate synagogues visible from the public street. Synagogue architecture blossomed. Large Jewish communities wished to show not only their wealth but also their newly acquired status as citizens by constructing magnificent synagogues. Handsome nineteenth synagogues form the period of Jewish imagination stand in virtually every country where there were Jewish communities. Most were built in revival styles then in fashion. Thus there were Neoclassical, Neo-Byzantine, Romanesque Revival Moorish Revival, Gothic Revival, and Greek Revival. There are Egyptian Revival synagogues and even one Mayan Revival synagogue. In the nineteenth and early twentieth century heyday of historicist architecture, however, most historicist synagogues, even the most magnificent ones, did not attempt a pure style, or even any particular style, and are best described as eclectic.
Chabad Lubavitch has made a practice of designing some of its Chabad Houses and centers as replicas of or homages to the architecture of 770 Eastern Parkway.
Central Europe: Polish?Lithuanian Commonwealth.
Wolpa Synagogue, 1920, Poland
The great exceptions to the rule that synagogues are built in the prevailing style of their time and place are the wooden synagogues of the former Polish?Lithuanian Commonwealth. They were a unique Jewish artistic and architectural form. Characteristic features include the independence of the pitched roof from the design of the interior domed ceiling. Elaborately carved, painted, domed, balconied and vaulted interiors. The architectural interest of the exterior lay in the large scale of the buildings, the multiple, horizontal lines of the tiered roofs, and the carved corbels that supported them. Wooden synagogues featured a single, large hall. In contrast to contemporary churches, there was no apse. Moreover, while contemporary churches featured imposing vestibules, the entry porches of the wooden synagogues was a low annex, usually with a simple lean-to roof. In these synagogues, the emphasis was on constructing a single, large, high-domed worship space.
According to art historian Stephen S. Kayser, the wooden synagogues of Poland with their painted and carved interiors were "a truly original and organic manifestation of artistic expression?the only real Jewish folk art in history."
According to Louis Lozowick, writing in 1947, the wooden synagogues were unique because, unlike all previous synagogues, they were not built in the architectural style of their region and era, but in a newly evolved and uniquely Jewish style, making them "a truly original folk expression," whose "originality does not lie alone in the exterior architecture, it lies equally in the beautiful and intricate wood carving of the interior."
Moreover, while in many parts of the world Jews were proscribed from entering the building trades and even from practicing the decorative arts of painting and woodcarving, the wooden synagogues were actually built by Jewish craftsmen.
Art historian Ori Z. Soltes points out that the wooden synagogues, unusual for that period in being large, identifiably Jewish buildings not hidden in courtyards or behind walls, were built not only during a Jewish "intellectual golden age" but in a time and place where "the local Jewish population was equal to or even greater than the Christian population.
Egyptian Revival.
Egyptian door, Hobart Synagogue
Egyptian Revival style synagogues were popular in the early nineteenth century. Rachel Wischnitzer argues that they were part of the fashion for Egyptian style inspired by Napoleon's invasion of Europe. According to Carol Herselle Krinsky, they were meant as imitations of the Temple of Solomon and intended by architects and governments to insult Jews by portraying Judaism as a primitive faith. According to Diana Muir Appelbaum, they were expressions of Jewish identity intended to advertise Jewish origins in ancient Israel.
Moorish influence.
Interior of Santa María la Blanca
In medieval Spain (both Al-Andalus and the Christian kingdoms), a host of synagogues were built, and it was usual to commission them from Moorish and later Mudéjar architects. Very few of these medieval synagogues, built with Moorish techniques and style, are conserved. The two best known Spanish synagogues are in Toledo, one known as El Tránsito, the other as Santa María la Blanca, and both, undoubtedly very beautiful, are now preserved as national monuments. The former is a small building containing very rich decorations; the latter is especially noteworthy. It is based upon Almohad style and contains long rows of octagonal columns with curiously carved capitals, from which spring Moorish arches supporting the roof.
Moorish Revival Great Synagogue of Plze?. Another significative Mudéjar synagogue is the one at Córdoba built in 1315. As in El Tránsito, the vegetal and geometrical stucco decorations are purely Moorish, but unlike the former, the epigraphic texts are in Hebrew.
After the expulsion from Spain there was a general feeling among wealthy Sephardim that Moorish architecture was appropriate in synagogues. By the mid-19th century, the style was adopted by the Ashkenazim of Central and Eastern Europe, who associated Moorish and Mudéjar architectural forms with the golden age of Jewry in Al-Andalus. As a consequence, Moorish Revival spread around the globe as a preferred style of synagogue architecture, although Moorish architecture is by no means Jewish, either in fact or in feeling. The Alhambra has furnished inspiration for innumerable synagogues, but seldom have its graceful proportions or its delicate modeling and elaborate ornamentation been successfully copied.
Modern synagogue architecture
In the modern period, synagogues have continued to be built in every popular architectural style, including Art Nouveau, Art Deco, International style, and all contemporary styles. In the post-World War II period "a period of post-war modernism," came to the fore, "characterized by assertive architectural gestures that had the strength and integrity to stand alone, without applied artwork or Jewish iconography." A notable work of Art Nouveau, pre-World War I Hungarian synagogue architecture is Budapest's Kazinczy Street Synagogue.
The most common general plan for the interior of the synagogue is an Ark at the eastern end opposite the entrance, and with an almemar or pulpit. In older or Orthodox synagogues with separate seating, there may be benches for the men on either side, and a women's gallery reached by staircases from the outer vestibule. Variations of this simple plan abound: the vestibule became larger, and the staircases to the women's gallery were separated from the vestibule and given more importance. As the buildings became larger, rows of columns were required to support the roof, but in every case the basilican form was retained. The Ark, formerly allowed a mere niche in the wall, was developed into the main architectural feature of the interior, and was flanked with columns, covered with a canopy and richly decorated. The almemar in many cases was joined to the platform in front of the Ark, and elaborate arrangements of steps were provided.
The Ark.
The Torah Ark (usually called Aron Hakodesh or Hekhál) is the most important feature of the interior, and is generally dignified by proper decoration and raised upon a suitable platform, reached by at least three steps, but often by more. It is usually crowned by the Ten Commandments. The position of the pulpit varies; it may be placed on either side of the Ark, and is occasionally found in the center of the steps.
Other interior arrangements.
The modern synagogue, besides containing the minister's study, trustees' rooms, choir-rooms, and organ-loft, devotes much space to school purposes; generally the entire lower floor is used for class-rooms. The interior treatment of the synagogue allows great latitude in design.
For the thirty-three synagogues of India, American architect and professor of architecture Jay A. Waronker has learned that these buildings tend to follow the Sephardic traditions of the tevah (or bimah, the raised platform where the service is led and Torah read) being freestanding and roughly in the middle of the sanctuary and the ark (called the hekhal by Sephardim and the aron ha-kodesh by Ashkenazim) engaged along the wall that is closest to Jerusalem. The hekhals are essentially cabinets or armoires storing the sefer Torahs. Seating, in the form of long wooden benches, is grouped around and facing the tevah. Men sit together on the main level of the sanctuary while women sit in a dedicated zone on the same level in the smaller synagogues or upstairs in a women's gallery.
Interesting architectural and planning exceptions to this common Sephardic formula are the Cochin synagogues in Kerala of far southwestern India. Here, on the gallery level and adjacent to the space provided for women and overlooking the sanctuary below, is a second tevah. This tevah was used for holidays and unique occasions. It is therefore interesting, on more special events, the woman are closest to the point where the religious service is being led.
In Baghdadi synagogues of India, the hekhals appear to be standard-sized cabinets from the outside (the side facing......
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68.Ants.
Ants are eusocial insects of the family Formicidae /f?r?m?s?di?/ and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from wasp-like ancestors in the mid-Cretaceous period between 110 and 130 million years ago and diversified after the rise of flowering plants. More than 12,500 of an estimated total of 22,000 species have been classified. They are easily identified by their elbowed antennae and the distinctive node-like structure that forms their slender waists.
Ants form colonies that range in size from a few dozen predatory individuals living in small natural cavities to highly organised colonies that may occupy large territories and consist of millions of individuals. Larger colonies consist mostly of sterile, wingless females forming castes of "workers", "soldiers", or other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called "queens". The colonies are described as superorganisms because the ants appear to operate as a unified entity, collectively working together to support the colony.
Ants have colonised almost every landmass on Earth. The only places lacking indigenous ants are Antarctica and a few remote or inhospitable islands. Ants thrive in most ecosystems and may form 15?25% of the terrestrial animal biomass. Their success in so many environments has been attributed to their social organisation and their ability to modify habitats, tap resources, and defend themselves. Their long co-evolution with other species has led to mimetic, commensal, parasitic, and mutualistic relationships.
Ant societies have division of labour, communication between individuals, and an ability to solve complex problems. These parallels with human societies have long been an inspiration and subject of study. Many human cultures make use of ants in cuisine, medication, and rituals. Some species are valued in their role as biological pest control agents. Their ability to exploit resources may bring ants into conflict with humans, however, as they can damage crops and invade buildings. Some species, such as the red imported fire ant (Solenopsis invicta), are regarded as invasive species, establishing themselves in areas where they have been introduced accidentally.
The word "ant" is derived from ante, emete of Middle English which are derived from ?mette of Old English, and is related to the dialectal Dutch emt and the Old High German ?meiza, hence the modern German Ameise. All of these words come from West Germanic *?maitij?, and the original meaning of the word was "the biter" (from Proto-Germanic *ai-, "off, away" + *mait- "cut"). The family name Formicidae is derived from the Latin form?ca ("ant") from which the words in other Romance languages, such as the Portuguese formiga, Italian formica, Spanish hormiga, Romanian furnic?, and French fourmi are derived. It has been hypothesised that a Proto-Indo-European word *morwi- was used, cf. Sanskrit vamrah, Latin form?ca, Greek ?????? mýrm?x, Old Church Slavonic mraviji, Old Irish moirb, Old Norse maurr, Dutch mier.
The family Formicidae belongs to the order Hymenoptera, which also includes sawflies, bees, and wasps. Ants evolved from a lineage within the aculeate wasps, and a 2013 study suggests they are a sister group of the Apoidea. In 1966, E. O. Wilson and his colleagues identified the fossil remains of an ant (Sphecomyrma) that lived in the Cretaceous period. The specimen, trapped in amber dating back to around 92 million years ago, has features found in some wasps, but not found in modern ants. Sphecomyrma possibly was a ground forager, while Haidomyrmex and Haidomyrmodes, related genera in subfamily Sphecomyrminae, are reconstructed as active arboreal predators. After the rise of flowering plants about 100 million years ago they diversified and assumed ecological dominance around 60 million years ago. Some groups, such as the Leptanillinae and Martialinae, are suggested to have diversified from early primitive ants that were likely to have been predators underneath the surface of the soil.
During the Cretaceous period, a few species of primitive ants ranged widely on the Laurasian supercontinent (the Northern Hemisphere). They were scarce in comparison to the populations of other insects, representing only about 1% of the entire insect population. Ants became dominant after adaptive radiation at the beginning of the Paleogene period. By the Oligocene and Miocene, ants had come to represent 20?40% of all insects found in major fossil deposits. Of the species that lived in the Eocene epoch, around one in 10 genera survive to the present. Genera surviving today comprise 56% of the genera in Baltic amber fossils (early Oligocene), and 92% of the genera in Dominican amber fossils (apparently early Miocene).
Termites, although sometimes called 'white ants', are not ants. They belong to the sub-order Isoptera within order Blattodea. Termites are more closely related to cockroaches and mantids. Termites are eusocial, but differ greatly in the genetics of reproduction. The similarity of their social structure to that of ants is attributed to convergent evolution. Velvet ants look like large ants, but are wingless female wasps.
Ants are found on all continents except Antarctica, and only a few large islands, such as Greenland, Iceland, parts of Polynesia and the Hawaiian Islands lack native ant species. Ants occupy a wide range of ecological niches, and are able to exploit a wide range of food resources either as direct or indirect herbivores, predators, and scavengers. Most species are omnivorous generalists, but a few are specialist feeders. Their ecological dominance may be measured by their biomass and estimates in different environments suggest that they contribute 15?20% (on average and nearly 25% in the tropics) of the total terrestrial animal biomass, which exceeds that of the vertebrates.
Ants range in size from 0.75 to 52 millimetres (0.030?2.0 in), the largest species being the fossil Titanomyrma giganteum, the queen of which was 6 centimetres (2.4 in) long with a wingspan of 15 centimetres (5.9 in). Ants vary in colour; most ants are red or black, but a few species are green and some tropical species have a metallic lustre. More than 12,000 species are currently known (with upper estimates of the potential existence of about 22,000), with the greatest diversity in the tropics. Taxonomic studies continue to resolve the classification and systematics of ants. Online databases of ant species, including AntBase and the Hymenoptera Name Server, help to keep track of the known and newly described species. The relative ease with which ants may be sampled and studied in ecosystems has made them useful as indicator species in biodiversity studies.
Ants are distinct in their morphology from other insects in having elbowed antennae, metapleural glands, and a strong constriction of their second abdominal segment into a node-like petiole. The head, mesosoma, and metasoma are the three distinct body segments. The petiole forms a narrow waist between their mesosoma (thorax plus the first abdominal segment, which is fused to it) and gaster (abdomen less the abdominal segments in the petiole). The petiole may be formed by one or two nodes (the second alone, or the second and third abdominal segments).
Like other insects, ants have an exoskeleton, an external covering that provides a protective casing around the body and a point of attachment for muscles, in contrast to the internal skeletons of humans and other vertebrates. Insects do not have lungs; oxygen and other gases such as carbon dioxide pass through their exoskeleton via tiny valves called spiracles. Insects also lack closed blood vessels; instead, they have a long, thin, perforated tube along the top of the body (called the "dorsal aorta") that functions like a heart, and pumps haemolymph toward the head, thus driving the circulation of the internal fluids. The nervous system consists of a ventral nerve cord that runs the length of the body, with several ganglia and branches along the way reaching into the extremities of the appendages.
Head.
An ant's head contains many sensory organs. Like most insects, ants have compound eyes made from numerous tiny lenses attached together. Ant eyes are good for acute movement detection, but do not offer a high resolution image. They also have three small ocelli (simple eyes) on the top of the head that detect light levels and polarization. Compared to vertebrates, most ants have poor-to-mediocre eyesight and a few subterranean species are completely blind. Some ants such as Australia's bulldog ant, however, have excellent vision and are capable of discriminating the distance and size of objects moving nearly a metre away.
Two antennae ("feelers") are attached to the head; these organs detect chemicals, air currents, and vibrations; they also are used to transmit and receive signals through touch. The head has two strong jaws, the mandibles, used to carry food, manipulate objects, construct nests, and for defence. In some species a small pocket (infrabuccal chamber) inside the mouth stores food, so it may be passed to other ants or their larvae.
Legs.
All six legs are attached to the mesosoma ("thorax") and terminate in a hooked claw.
Wings.
Only reproductive ants, queens and males, have wings. Queens shed the wings after the nuptial flight, leaving visible stubs, a distinguishing feature of queens. Wingless queens (ergatoids) and males occur in a few species, however.
Metasoma.
The metasoma (the "abdomen") of the ant houses important internal organs, including those of the reproductive, respiratory (tracheae), and excretory systems. Workers of many species have their egg-laying structures modified into stings that are used for subduing prey and defending their nests.
Polymorphism.
Seven Leafcutter ant workers of various castes (left) and two Queens (right)
In the colonies of a few ant species, there are physical castes?workers in distinct size-classes, called minor, median, and major workers. Often, the larger ants have disproportionately larger heads, and correspondingly stronger mandibles. Such individuals are sometimes called "soldier" ants because their stronger mandibles make them more effective in fighting, although they still are workers and their "duties" typically do not vary greatly from the minor or median workers. In a few species, the median workers are absent, creating a sharp divide between the minors and majors. Weaver ants, for example, have a distinct bimodal size distribution. Some other species show continuous variation in the size of workers. The smallest and largest workers in Pheidologeton diversus show nearly a 500-fold difference in their dry-weights. Workers cannot mate; however, because of the haplodiploid sex-determination system in ants, workers of a number of species can lay unfertilised eggs that become fully fertile, haploid males. The role of workers may change with their age and in some species, such as honeypot ants, young workers are fed until their gasters are distended, and act as living food storage vessels. These food storage workers are called repletes. For instance, these replete workers develop in the North American honeypot ant Myrmecocystus mexicanus. Rissing found that usually the largest workers in the colony develop into repletes, and if repletes are removed from the colony other workers become repletes, demonstrating the flexibility of this particular polymorphism. This polymorphism in morphology and behaviour of workers initially was thought to be determined by environmental factors such as nutrition and hormones that led to different developmental paths; however, genetic differences between worker castes have been noted in Acromyrmex sp. These polymorphisms are caused by relatively small genetic changes; differences in a single gene of Solenopsis invicta can decide whether the colony will have single or multiple queens. The Australian jack jumper ant (Myrmecia pilosula) has only a single pair of chromosomes (with the males having just one chromosome as they are haploid), the lowest number known for any animal, making it an interesting subject for studies in the genetics and developmental biology of social insects.
The life of an ant starts from an egg. If the egg is fertilised, the progeny will be female (diploid); if not, it will be male (haploid). Ants develop by complete metamorphosis with the larva stages passing through a pupal stage before emerging as an adult. The larva is largely immobile and is fed and cared for by workers. Food is given to the larvae by trophallaxis, a process in which an ant regurgitates liquid food held in its crop. This is also how adults share food, stored in the "social stomach". Larvae, especially in the later stages, may also be provided solid food such as trophic eggs, pieces of prey, and seeds brought by workers.
The larvae grow through a series of four or five moults and enter the pupal stage. The pupa has the appendages free and not fused to the body as in a butterfly pupa. The differentiation into queens and workers (which are both female), and different castes of workers (when they exist), is influenced in some species by the nutrition the larvae obtain. Genetic influences and the control of gene expression by the developmental environment are complex and the determination of caste continues to be a subject of research. Larvae and pupae need to be kept at fairly constant temperatures to ensure proper development, and so often, are moved around among the various brood chambers within the colony.
A new worker spends the first few days of its adult life caring for the queen and young. She then graduates to digging and other nest work, and later to defending the nest and foraging. These changes are sometimes fairly sudden, and define what are called temporal castes. An explanation for the sequence is suggested by the high casualties involved in foraging, making it an acceptable risk only for ants who are older and are likely to die soon of natural causes.
Most ant species have a system in which only the queen and breeding females have the ability to mate. Contrary to popular belief, some ant nests have multiple queens, while others may exist without queens. Workers with the ability to reproduce are called "gamergates" and colonies that lack queens are then called gamergate colonies; colonies with queens are said to be queen-right. The winged male ants, called drones, emerge from pupae along with the breeding females (although some species, such as army ants, have wingless queens), and do nothing in life except eat and mate.
Most ants are univoltine, producing a new generation each year. During the species-specific breeding period, new reproductives, females and winged males leave the colony in what is called a nuptial flight. Typically, the males take flight before the females. Males then use visual cues to find a common mating ground, for example, a landmark such as a pine tree to which other males in the area converge. Males secrete a mating pheromone that females follow. Females of some species mate with just one male, but in others they may mate with as many as ten or more different males.
Mated females then seek a suitable place to begin a colony. There, they break off their wings and begin to lay and care for eggs. The females store the sperm they obtain during their nuptial flight to selectively fertilise future eggs. The first workers to hatch are weak and smaller than later workers, but they begin to serve the colony immediately. They enlarge the nest, forage for food, and care for the other eggs. This is how new colonies start in most ant species. Species that have multiple queens may have a queen leaving the nest along with some workers to found a colony at a new site, a process akin to swarming in honeybees.
Ants mating.
A wide range of reproductive strategies have been noted in ant species. Females of many species are known to be capable of reproducing asexually through thelytokous parthenogenesis.
Ant colonies can be long-lived. The queens can live for up to 30 years, and workers live from 1 to 3 years. Males, however, are more transitory, being quite short-lived and surviving for only a few weeks. Ant queens are estimated to live 100 times longer than solitary insects of a similar size.
Ants are active all year long in the tropics, but, in cooler regions, they survive the winter in a state of dormancy or inactivity. The forms of inactivity are varied and some temperate species have larvae going into the inactive state, (diapause), while in others, the adults alone pass the winter in a state of reduced activity.
Communication.
Ants communicate with each other using pheromones, sounds, and touch. The use of pheromones as chemical signals is more developed in ants, such as the red harvester ant, than in other hymenopteran groups. Like other insects, ants perceive smells with their long, thin, and mobile antennae. The paired antennae provide information about the direction and intensity of scents. Since most ants live on the ground, they use the soil surface to leave pheromone trails that may be followed by other ants. In species that forage in groups, a forager that finds food marks a trail on the way back to the colony; this trail is followed by other ants, these ants then reinforce the trail when they head back with food to the colony. When the food source is exhausted, no new trails are marked by returning ants and the scent slowly dissipates. This behaviour helps ants deal with changes in their environment. For instance, when an established path to a food source is blocked by an obstacle, the foragers leave the path to explore new routes. If an ant is successful, it leaves a new trail marking the shortest route on its return. Successful trails are followed by more ants, reinforcing better routes and gradually identifying the best path.
Ants use pheromones for more than just making trails. A crushed ant emits an alarm pheromone that sends nearby ants into an attack frenzy and attracts more ants from farther away. Several ant species even use "propaganda pheromones" to confuse enemy ants and make them fight among themselves. Pheromones are produced by a wide range of structures including Dufour's glands, poison glands and glands on the hindgut, pygidium, rectum, sternum, and hind tibia. Pheromones also are exchanged, mixed with food, and passed by trophallaxis, transferring information within the colony. This allows other ants to detect what task group (e.g., foraging or nest maintenance) other colony members belong to. In ant species with queen castes, when the dominant queen stops producing a specific pheromone, workers begin to raise new queens in the colony.
Some ants produce sounds by stridulation, using the gaster segments and their mandibles. Sounds may be used to communicate with colony members or with other species.
Defence.
A Plectroctena sp. attacks another of its kind to protect its territory
See also Insect defences
Ants attack and defend themselves by biting and, in many species, by stinging, often injecting or spraying chemicals, such as formic acid in the case of formicine ants, alkaloids and piperidines in fire ants, and a variety of protein components in other ants. Bullet ants (Paraponera), located in Central and South America, are considered to have the most painful sting of any insect, although it is usually not fatal to humans. This sting is given the highest rating on the Schmidt Sting Pain Index.
The sting of jack jumper ants can be fatal, and an antivenom has been developed for it.
Fire ants, Solenopsis spp., are unique in having a poison sac containing piperidine alkaloids. Their stings are painful and can be dangerous to hypersensitive people.
A weaver ant in fighting position, mandibles wide open
Trap-jaw ants of the genus Odontomachus are equipped with mandibles called trap-jaws, which snap shut faster than any other predatory appendages within the animal kingdom. One study of Odontomachus bauri recorded peak speeds of between 126 and 230 km/h (78 ? 143 mph), with the jaws closing within 130 microseconds on average. The ants were also observed to use their jaws as a catapult to eject intruders or fling themselves backward to escape a threat. Before striking, the ant opens its mandibles extremely widely and locks them in this position by an internal mechanism. Energy is stored in a thick band of muscle and explosively released when triggered by the stimulation of sensory organs resembling hairs on the inside of the mandibles. The mandibles also permit slow and fine movements for other tasks. Trap-jaws also are seen in the following genera: Anochetus, Orectognathus, and Strumigenys, plus some members of the Dacetini tribe, which are viewed as examples of convergent evolution.
A Malaysian species of ant in the Camponotus cylindricus group has enlarged mandibular glands that extend into their gaster. When disturbed, workers rupture the membrane of the gaster, causing a burst of secretions containing acetophenones and other chemicals that immobilise small insect attackers. The worker subsequently dies.
Suicidal defences by workers are also noted in a Brazilian ant, Forelius pusillus, where a small group of ants leaves the security of the nest after sealing the entrance from the outside each evening.
Ant mound holes prevent water from entering the nest during rain
In addition to defence against predators, ants need to protect their colonies from pathogens. Some worker ants maintain the hygiene of the colony and their activities include undertaking or necrophory, the disposal of dead nest-mates. Oleic acid has been identified as the compound released from dead ants that triggers necrophoric behaviour in Atta mexicana while workers of Linepithema humile react to the absence of characteristic chemicals (dolichodial and iridomyrmecin) present on the cuticle of their living nestmates to trigger similar behaviour.
Nests may be protected from physical threats such as flooding and overheating by elaborate nest architecture. Workers of Cataulacus muticus, an arboreal species that lives in plant hollows, respond to flooding by drinking water inside the nest, and excreting it outside. Camponotus anderseni, which nests in the cavities of wood in mangrove habitats, deals with submergence under water by switching to anaerobic respiration.
Learning.
Many animals can learn behaviours by imitation, but ants may be the only group apart from mammals where interactive teaching has been observed. A knowledgeable forager of Temnothorax albipennis will lead a naive nest-mate to newly discovered food by the process of tandem running. The follower obtains knowledge through its leading tutor. The leader is acutely sensitive to the progress of the follower and slows down when the follower lags and speeds up when the follower gets too close.
Controlled experiments with colonies of Cerapachys biroi suggest that an individual may choose nest roles based on her previous experience. An entire generation of identical workers was divided into two groups whose outcome in food foraging was controlled. One group was continually rewarded with prey, while it was made certain that the other failed. As a result, members of the successful group intensified their foraging attempts while the unsuccessful group ventured out fewer and fewer times. A month later, the successful foragers continued in their role while the others had moved to specialise in brood care.
Nest construction.
Complex nests are built by many ant species, but other species are nomadic and do not build permanent structures. Ants may form subterranean nests or build them on trees. These nests may be found in the ground, under stones or logs, inside logs, hollow stems, or even acorns. The materials used for construction include soil and plant matter, and ants carefully select their nest sites; Temnothorax albipennis will avoid sites with dead ants, as these may indicate the presence of pests or disease. They are quick to abandon established nests at the first sign of threats.
The army ants of South America, such as the Eciton burchellii species, and the driver ants of Africa do not build permanent nests, but instead, alternate between nomadism and stages where the workers form a temporary nest (bivouac) from their own bodies, by holding each other together.
Weaver ant (Oecophylla spp.) workers build nests in trees by attaching leaves together, first pulling them together with bridges of workers and then inducing their larvae to produce silk as they are moved along the leaf edges. Similar forms of nest construction are seen in some species of Polyrhachis.
Formica polyctena, among other ant species, constructs nests that maintain a relatively constant interior temperature that aids in the development of larvae. The ants maintain the nest temperature by choosing the location, nest materials, controlling ventilation and maintaining the heat from solar radiation, worker activity and metabolism, and in some moist nests, microbial activity in the nest materials.
Some ant species, such as those that use natural cavities, can be opportunistic and make use of the controlled micro-climate provided inside human dwellings and other artificial structures to house their colonies and nest structures.
Cultivation of food.
Myrmecocystus, honeypot ants, store food to prevent colony famine
Most ants are generalist predators, scavengers, and indirect herbivores, but a few have evolved specialised ways of obtaining nutrition. It is believed that many ant species that engage in indirect herbivory rely on specialized symbiosis with their gut microbes to upgrade the nutritional value of the food they collect and allow them to survive in nitrogen poor regions, such as rainforrest canopies. Leafcutter ants (Atta and Acromyrmex) feed exclusively on a fungus that grows only within their colonies. They continually collect leaves which are taken to the colony, cut into tiny pieces and placed in fungal gardens. Workers specialise in related tasks according to their sizes. The largest ants cut stalks, smaller workers chew the leaves and the smallest tend the fungus. Leafcutter ants are sensitive enough to recognise the reaction of the fungus to different plant material, apparently detecting chemical signals from the fungus. If a particular type of leaf is found to be toxic to the fungus, the colony will no longer collect it. The ants feed on structures produced by the fungi called gongylidia. Symbiotic bacteria on the exterior surface of the ants produce antibiotics that kill bacteria introduced into the nest that may harm the fungi.
Navigation.
Foraging ants travel distances of up to 200 metres (700 ft) from their nest and scent trails allow them to find their way back even in the dark. In hot and arid regions, day-foraging ants face death by desiccation, so the ability to find the shortest route back to the nest reduces that risk. Diurnal desert ants of the genus Cataglyphis such as the Sahara desert ant navigate by keeping track of direction as well as distance travelled. Distances travelled are measured using an internal pedometer that keeps count of the steps taken and also by evaluating the movement of objects in their visual field (optical flow). Directions are measured using the position of the sun. They integrate this information to find the shortest route back to their nest. Like all ants, they can also make use of visual landmarks when available as well as olfactory and tactile cues to navigate. Some species of ant are able to use the Earth's magnetic field for navigation. The compound eyes of ants have specialised cells that detect polarised light from the Sun, which is used to determine direction. These polarization detectors are sensitive in the ultraviolet region of the light spectrum. In some army ant species, a group of foragers who become separated from the main column sometimes may turn back on themselves and form a circular ant mill. The workers may then run around continuously until they die of exhaustion.
Locomotion.
The female worker ants do not have wings and reproductive females lose their wings after their mating flights in order to begin their colonies. Therefore, unlike their wasp ancestors, most ants travel by walking. Some species are capable of leaping. For example, Jerdon's jumping ant (Harpegnathos saltator) is able to jump by synchronising the action of its mid and hind pairs of legs. There are several species of gliding ant including Cephalotes atratus; this may be a common trait among most arboreal ants. Ants with this ability are able to control the direction of their descent while falling.
Other species of ants can form chains to bridge gaps over water, underground, or through spaces in vegetation. Some species also form floating rafts that help them survive floods. These rafts may also have a role in allowing ants to colonise islands. Polyrhachis sokolova, a species of ant found in Australian mangrove swamps, can swim and live in underwater nests. Since they lack gills, they go to trapped pockets of air in the submerged nests to breathe.
Cooperation and competition.
Meat-eater ants feeding on a cicada, social ants cooperate and collectively gather food
Not all ants have the same kind of societies. The Australian bulldog ants are among the biggest and most basal of ants. Like virtually all ants, they are eusocial, but their social behaviour is poorly developed compared to other species. Each individual hunts alone, using her large eyes instead of chemical senses to find prey.
Some species (such as Tetramorium caespitum) attack and take over neighbouring ant colonies. Others are less expansionist, but just as aggressive; they invade colonies to steal eggs or larvae, which they either eat or raise as workers or slaves. Extreme specialists among these slave-raiding ants, such as the Amazon ants, are incapable of feeding themselves and need captured workers to survive. Captured workers of the enslaved species Temnothorax have evolved a counter strategy, destroying just the female pupae of the slave-making Protomognathus americanus, but sparing the males (who don't take part in slave-raiding as adults).
A worker Harpegnathos saltator (a jumping ant) engaged in battle with a rival colony's queen
Ants identify kin and nestmates through their scent, which comes from hydrocarbon-laced secretions that coat their exoskeletons. If an ant is separated from its original colony, it will eventually lose the colony scent. Any ant that enters a colony without a matching scent will be attacked. Also, the reason why two separate colonies of ants will attack each other even if they are of the same species is because the genes responsible for pheromone production are different between them. The Argentine ant, however, does not have this characteristic, due to lack of genetic diversity, and has become a global pest because of it.
Parasitic ant species enter the colonies of host ants and establish themselves as social parasites; species such as Strumigenys xenos are entirely parasitic and do not have workers, but instead, rely on the food gathered by their Strumigenys perplexa hosts. This form of parasitism is seen across many ant genera, but the parasitic ant is usually a species that is closely related to its host. A variety of methods are employed to enter the nest of the host ant. A parasitic queen may enter the host nest before the first brood has hatched, establishing herself prior to development of a colony scent. Other species use pheromones to confuse the host ants or to trick them into carrying the parasitic queen into the nest. Some simply fight their way into the nest.
A conflict between the sexes of a species is seen in some species of ants with these reproducers apparently competing to produce offspring that are as closely related to them as possible. The most extreme form involves the production of clonal offspring. An extreme of sexual conflict is seen in Wasmannia auropunctata, where the queens produce diploid daughters by thelytokous parthenogenesis and males produce clones by a process whereby a diploid egg loses its maternal contribution to produce haploid males who are clones of the father.
Relationships with other organisms.
The spider Myrmarachne plataleoides (female shown) mimics weaver ants to avoid predators.
Ants form symbiotic associations with a range of species, including other ant species, other insects, plants, and fungi. They also are preyed on by many animals and even certain fungi. Some arthropod species spend part of their lives within ant nests, either preying on ants, their larvae, and eggs, consuming the food stores of the ants, or avoiding predators. These inquilines may bear a close resemblance to ants. The nature of this ant mimicry (myrmecomorphy) varies, with some cases involving Batesian mimicry, where the mimic reduces the risk of predation. Others show Wasmannian mimicry, a form of mimicry seen only in inquilines.
Aphids and other hemipteran insects secrete a sweet liquid called honeydew, when they feed on plant sap. The sugars in honeydew are a high-energy food source, which many ant species collect. In some cases, the aphids secrete the honeydew in response to ants tapping them with their antennae. The ants in turn keep predators away from the aphids and will move them from one feeding location to another. When migrating to a new area, many colonies will take the aphids with them, to ensure a continued supply of honeydew. Ants also tend mealybugs to harvest their honeydew. Mealybugs may become a serious pest of pineapples if ants are present to protect mealybugs from their natural enemies.
Myrmecophilous (ant-loving) caterpillars of the butterfly family Lycaenidae (e.g., blues, coppers, or hairstreaks) are herded by the ants, led to feeding areas in the daytime, and brought inside the ants' nest at night. The caterpillars have a gland which secretes honeydew when the ants massage them. Some caterpillars produce vibrations and sounds that are perceived by the ants. Other caterpillars have evolved from ant-loving to ant-eating: these myrmecophagous caterpillars secrete a pheromone that makes the ants act as if the caterpillar is one of their own larvae. The caterpillar is then taken into the ant nest where it feeds on the ant larvae. Fungus-growing ants that make up the tribe Attini, including leafcutter ants, cultivate certain species of fungus in the Leucoagaricus or Leucocoprinus genera of the Agaricaceae family. In this ant-fungus mutualism, both species depend on each other for survival. The ant Allomerus decemarticulatus has evolved a three-way association with the host plant, Hirtella physophora (Chrysobalanaceae), and a sticky fungus which is used to trap their insect prey.
Ants may obtain nectar from flowers such as the dandelion but are only rarely known to pollinate flowers.
Lemon ants make devil's gardens by killing surrounding plants with their stings and leaving a pure patch of lemon ant trees, (Duroia hirsuta). This modification of the forest provides the ants with more nesting sites inside the stems of the Duroia trees. Although some ants obtain nectar from flowers, pollination by ants is somewhat rare. Some plants have special nectar exuding structures, extrafloral nectaries that provide food for ants, which in turn protect the plant from more damaging herbivorous insects. Species such as the bullhorn acacia (Acacia cornigera) in Central America have hollow thorns that house colonies of stinging ants (Pseudomyrmex ferruginea) who defend the tree against insects, browsing mammals, and epiphytic vines. Isotopic labelling studies suggest that plants also obtain nitrogen from the ants. In return, the ants obtain food from protein- and lipid-rich Beltian bodies. Another example of this type of ectosymbiosis comes from the Macaranga tree, which has stems adapted to house colonies of Crematogaster ants.
Many tropical tree species have seeds that are dispersed by ants. Seed dispersal by ants or myrmecochory is widespread and new estimates suggest that nearly 9% of all plant species may have such ant associations. Some plants in fire-prone grassland systems are particularly dependent on ants for their survival and dispersal as the seeds are transported to safety below the ground. Many ant-dispersed seeds have special external structures, elaiosomes, that are sought after by ants as food.
A convergence, possibly a form of mimicry, is seen in the eggs of stick insects. They have an edible elaiosome-like structure and are taken into the ant nest where the young hatch.
Most ants are predatory and some prey on and obtain food from other social insects including other ants. Some species specialise in preying on termites (Megaponera and Termitopone) while a few Cerapachyinae prey on other ants. Some termites, including Nasutitermes corniger, form associations with certain ant species to keep away predatory ant species. The tropical wasp Mischocyttarus drewseni coats the pedicel of its nest with an ant-repellent chemical. It is suggested that many tropical wasps may build their nests in trees and cover them to protect themselves from ants. Other wasps such as A. multipicta defend against ants by blasting them off the nest with bursts of wing buzzing. Stingless bees (Trigona and Melipona) use chemical defences against ants. Certain species of ants have the power to drive certain wasps, such as Polybia occidentalis to extinction if they attack more than once and the wasps cannot keep up with rebuilding their nest.
Flies in the Old World genus Bengalia (Calliphoridae) prey on ants and are kleptoparasites, snatching prey or brood from the mandibles of adult ants. Wingless and legless females of the Malaysian phorid fly (Vestigipoda myrmolarvoidea) live in the nests of ants of the genus Aenictus and are cared for by the ants.
Fungi in the genera Cordyceps and Ophiocordyceps infect ants. Ants react to their infection by climbing up plants and sinking their mandibles into plant tissue. The fungus kills the ants, grows on their remains, and produces a fruiting body. It appears that the fungus alters the behaviour of the ant to help disperse its spores in a microhabitat that best suits the fungus. Strepsipteran parasites also manipulate their ant host to climb grass stems, to help the parasite find mates.
A nematode (Myrmeconema neotropicum) that infects canopy ants (Cephalotes atratus) causes the black-coloured gasters of workers to turn red. The parasite also alters the behaviour of the ant, causing them to carry their gasters high. The conspicuous red gasters are mistaken by birds for ripe fruits, such as Hyeronima alchorneoides, and eaten. The droppings of the bird are collected by other ants and fed to their young, leading to further spread of the nematode.
South American poison dart frogs in the genus Dendrobates feed mainly on ants, and the toxins in their skin may come from the ants.
Army ants forage in a wide roving column, attacking any animals in that path that are unable to escape. In Central and South America, Eciton burchellii is the swarming ant most commonly attended by "ant-following" birds such as antbirds and woodcreepers. This behaviour was once considered mutualistic, but later studies found the birds to be parasitic. Although direct kleptoparasitism (birds stealing food from the ants' grasp) is rare, the birds eat many prey insects that the ants would otherwise eat and thus decrease their foraging success. Birds indulge in a peculiar behaviour called anting that, as yet, is not fully understood. Here birds rest on ant nests, or pick and drop ants onto their wings and feathers; this may be a means to remove ectoparasites from the birds.
Anteaters, aardvarks, pangolins, echidnas and numbats have special adaptations for living on a diet of ants. These adaptations include long, sticky tongues to capture ants and strong claws to break into ant nests. Brown bears (Ursus arctos) have been found to feed on ants. About 12%, 16%, and 4% of their faecal volume in spring, summer, and autumn, respectively, is composed of ants.
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Inglish Site.15.
*
TO THE THRISE HO-
NOVRABLE AND EVER LY-
VING VERTVES OF SYR PHILLIP
SYDNEY KNIGHT, SYR JAMES JESUS SINGLETON, SYR CANARIS, SYR LAVRENTI BERIA ; AND TO THE
RIGHT HONORABLE AND OTHERS WHAT-
SOEVER, WHO LIVING LOVED THEM,
AND BEING DEAD GIVE THEM
THEIRE DVE.
***
In the beginning there is darkness. The screen erupts in blue, then a cascade of thick, white hexadecimal numbers and cracked language, ?UnusedStk? and ?AllocMem.? Black screen cedes to blue to white and a pair of scales appear, crossed by a sword, both images drawn in the jagged, bitmapped graphics of Windows 1.0-era clip-art?light grey and yellow on a background of light cyan. Blue text proclaims, ?God on tap!?
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Introduction.
Yes i am getting a little Mobi-Literate(ML) by experimenting literary on my Mobile Phone. Peoplecall it Typographical Laziness(TL).
The first accidental entries for the this part of this encyclopedia.
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This is TempleOS V2.17, the welcome screen explains, a ?Public Domain Operating System? produced by Trivial Solutions of Las Vegas, Nevada. It greets the user with a riot of 16-color, scrolling, blinking text; depending on your frame of reference, it might recall ?DESQview, the ?Commodore 64, or a host of early DOS-based graphical user interfaces. In style if not in specifics, it evokes a particular era, a time when the then-new concept of ?personal computing? necessarily meant programming and tinkering and breaking things.
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Index.
67.Synagogue
68.Ants.
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67.Synagogue:Maat, or divine order.
A synagogue, also spelled synagog (from Greek ????????, transliterated synagog?, meaning "assembly"; ??? ???? beth knesset, meaning "house of assembly"; ??? ????? beth t'fila, meaning "house of prayer"; ??? shul; ?????? esnoga; ??? kahal), is a Jewish house of prayer.
Synagogues have a large hall for prayer (the main sanctuary), and may also have smaller rooms for study and sometimes a social hall and offices. Some have a separate room for Torah study, called the beth midrash (Sefaradi/sefardim) "beis midrash (Ashkenazi/Ashkenazim)???? ???? ("House of Study").
Synagogues are consecrated spaces that can be used only for the purpose of prayer; however a synagogue is not necessary for worship. Communal Jewish worship can be carried out wherever ten Jews (a minyan) assemble. Worship can also be carried out alone or with fewer than ten people assembled together. However there are certain prayers that are communal prayers and therefore can be recited only by a minyan. The synagogue does not replace the long-since destroyed Temple in Jerusalem.
Israelis use the Hebrew term beyt knesset (assembly house). Jews of Ashkenazi descent have traditionally used the Yiddish term "shul" (cognate with the German Schule, school) in everyday speech. Spanish and Portuguese Jews call the synagogue an esnoga. Persian Jews and Karaite Jews use the term Kenesa, which is derived from Aramaic, and some Arabic-speaking Jews use knis. Some Reform and Conservative Jews use the word "temple". The Greek word "Synagogue" is a good all-around term, used in English (and German and French), to cover the preceding possibilities.
Although synagogues existed a long time before the destruction of the 2nd Temple in 70 CE, communal worship in the time while the Temple still stood centered around the korbanot ("sacrificial offerings") brought by the kohanim ("priests") in the Holy Temple. The all-day Yom Kippur service, in fact, was an event in which the congregation both observed the movements of the kohen gadol ("the high priest") as he offered the day's sacrifices and prayed for his success.
During the Babylonian captivity (586?537 BCE) the Men of the Great Assembly formalized and standardized the language of the Jewish prayers. Prior to that people prayed as they saw fit, with each individual praying in his or her own way, and there were no standard prayers that were recited. Rabbi Yohanan ben Zakkai, one of the leaders at the end of the Second Temple era, promulgated the idea of creating individual houses of worship in whatever locale Jews found themselves. This contributed to the continuity of the Jewish people by maintaining a unique identity and a portable way of worship despite the destruction of the Temple, according to many historians.
Synagogues in the sense of purpose-built spaces for worship, or rooms originally constructed for some other purpose but reserved for formal, communal prayer, however, existed long before the destruction of the Second Temple. The earliest archaeological evidence for the existence of very early synagogues comes from Egypt, where stone synagogue dedication inscriptions dating from the 3rd century BCE prove that synagogues existed by that date. A synagogue dating from between 75 and 50 BCE has been uncovered at a Hasmonean-era winter palace near Jericho. More than a dozen Second Temple era synagogues have been identified by archaeologists.
Any Jew or group of Jews can build a synagogue. Synagogues have been constructed by ancient Jewish kings, by wealthy patrons, as part of a wide range of human institutions including secular educational institutions, governments, and hotels, by the entire community of Jews living in a particular place, or by sub-groups of Jews arrayed according to occupation, ethnicity (i.e. the Sephardic, Polish or Persian Jews of a town), style of religious observance (i.e., a Reform or an Orthodox synagogue), or by the followers of a particular rabbi.
There is no set blueprint for synagogues and the architectural shapes and interior designs of synagogues vary greatly. In fact, the influence from other local religious buildings can often be seen in synagogue arches, domes and towers.
Historically, synagogues were built in the prevailing architectural style of their time and place. Thus, the synagogue in Kaifeng, China looked very like Chinese temples of that region and era, with its outer wall and open garden in which several buildings were arranged. The styles of the earliest synagogues resembled the temples of other sects of the eastern Roman Empire. The surviving synagogues of medieval Spain are embellished with mudéjar plasterwork. The surviving medieval synagogues in Budapest and Prague are typical Gothic structures.
The emancipation of Jews in European countries not only enabled Jews to enter fields of enterprise from which they were formerly barred, but gave them the right to build synagogues without needing special permissions, synagogue architecture blossomed. Large Jewish communities wished to show not only their wealth but also their newly acquired status as citizens by constructing magnificent synagogues. These were built across Europe and in the United States in all of the historicist or revival styles then in fashion. Thus there were Neoclassical, Neo-Byzantine, Romanesque Revival, Moorish Revival, Gothic Revival, and Greek Revival. There are Egyptian Revival synagogues and even one Mayan Revival synagogue. In the 19th century and early 20th century heyday of historicist architecture, however, most historicist synagogues, even the most magnificent ones, did not attempt a pure style, or even any particular style, and are best described as eclectic.
In the post-war era, synagogue architecture abandoned historicist styles for modernism.
All synagogues contain a bimah, a table from which the Torah is read, and a desk for the prayer leader;
The Torah ark, (Hebrew: Aron Kodesh????? ????) (called the heikhal????? [temple] by Sephardim) is a cabinet in which the Torah scrolls are kept.
The ark in a synagogue is almost always positioned in such a way such that those who face it are facing towards Jerusalem. Thus, sanctuary seating plans in the Western world generally face east, while those east of Israel face west. Sanctuaries in Israel face towards Jerusalem. Occasionally synagogues face other directions for structural reasons; in such cases, some individuals might turn to face Jerusalem when standing for prayers, but the congregation as a whole does not.
The ark is reminiscent of the Ark of the Covenant which held the tablets inscribed with the Ten Commandments. This is the holiest spot in a synagogue, equivalent to the Holy of Holies. The ark is often closed with an ornate curtain, the parochet ?????, which hangs outside or inside the ark doors.
A large, raised, reader's platform called the bimah (????) by Ashkenazim and tebah by Sephardim, where the Torah scroll is placed to be read is a feature of all synagogues. In Sephardi synagogues it is also used as the prayer leader's reading desk.
Other traditional features include a continually lit lamp or lantern, usually electric in contemporary synagogues, called the ner tamid (?? ????), the "Eternal Light", used as a reminder of the western lamp of the menorah of the Temple in Jerusalem, which remained miraculously lit perpetually. Many have an elaborate chair named for the prophet Elijah which is only sat upon during the ceremony of Brit milah. Many synagogues have a large seven-branched candelabrum commemorating the full Menorah. Most contemporary synagogues also feature a lectern for the rabbi.
A synagogue may be decorated with artwork, but in the Rabbinic and Orthodox tradition, three-dimensional sculptures and depictions of the human body are not allowed as these are considered akin to idolatry.
Until the 19th century, an Ashkenazi synagogue, all seats most often faced the 'Torah Ark. In a Sephardi synagogue, seats were usually arranged around the perimeter of the sanctuary, but when the worshipers stood up to pray, everyone faced the Ark. In Ashkenazi synagogues The Torah was read on a reader's table located in the center of the room, while the leader of the prayer service, the Hazzan, stood at his own lectern or table, facing the Ark. In Sephardic synagogues, the table for reading the Torah was commonly placed at the opposite side of the room from the Torah Ark, leaving the center of the floor empty for the use of a ceremonial procession carrying the Torah between the Ark and the reading table.
Denominational differences.
New York's Reform Temple Emanu-El
Orthodox synagogues feature a partition (mechitzah) dividing the men's and women's seating areas, or a separate women's section located on a balcony.
The German Reform movement which arose in the early 19th century made many changes to the traditional look of the synagogue, keeping with its desire to simultaneously stay Jewish yet be accepted by the host culture.
The first Reform synagogue, which opened in Hamburg in 1811, introduced changes that made the synagogue look more like a church. These included: the installation of an organ to accompany the prayers (even on Shabbat, when musical instruments are proscribed by halakha, a choir to accompany the Hazzan, and vestments for the synagogue rabbi to wear.
In following decades, the central reader's table, the Bimah, was moved to the front of the Reform sanctuary?previously unheard-of[citation needed] in Orthodox synagogues. The rabbi now delivered his sermon from the front, much as the Christian ministers delivered their sermons in a church. The synagogue was renamed a "temple", to emphasize that the movement no longer looked forward to the rebuilding of the Temple in Jerusalem.
Synagogue architecture often follows styles in vogue at the place and time of construction. There is no set blueprint for synagogues and the architectural shapes and interior designs of synagogues vary greatly. According to tradition, the Divine Presence (Shekhinah) can be found wherever there is a minyan, a quorum, of ten. A synagogue always contains an ark, called aron ha-kodesh by Ashkenazim and hekhal by Sephardim, where the Torah scrolls are kept.
The ark may be more or less elaborate, even a cabinet not structurally integral to the building or a portable arrangement whereby a Torah is brought into a space temporarily used for worship. There must also be a table from which the Torah is read. The table, called bimah by eastern Ashkenazim, almemmar (or balemmer) by Central and Western Ashkenazim and tebah by Sephardim, where the Torah is read (and from where the services are conducted in Sephardi synagogues) can range from an elaborate platform integral to the building (many early modern synagogues of central Europe featured bimahs with pillars that rose to support the ceiling), to elaborate free-standing raised platforms, to simple tables. A ner tamid, a constantly lit light as a reminder of the constantly lit menorah of the Temple in Jerusalem. Many synagogues, mainly in Ashkenazi communities, feature a pulpit facing the congregation from which to address the assembled. All synagogues require an amud (Hebrew for "post" or "column"), a desk facing the Ark from which the Hazzan (reader, or prayer leader) leads the prayers.
A synagogue may or may not have artwork, synagogues range from simple, unadorned prayer rooms to elaborately decorated buildings in every architectural style.
The synagogue, or if it is a multi-purpose building, prayer sanctuaries within the synagogue, should face towards Jerusalem. Thus sanctuaries in the Western world generally face east, while those east of Israel face west. Sanctuaries in Israel face towards Jerusalem. But this orientation need not be exact, and occasionally synagogues face other directions for structural reasons, in which case the community may face Jerusalem when standing for prayers.
There are only a few Talmudic instructions on how synagogues could be built, namely that they had to have windows and that they had to be taller than other buildings in town, synagogue plans were generally unrestricted, and even these few rules were often disregarded (especially as Jews were often not allowed to build tall buildings). There is no set blueprint for synagogues and the architectural shapes as well as interior designs of synagogues vary greatly.
Touro Synagogue, the oldest synagogue building still standing in the United States, built in Georgian style.
Baroque style W?odawa Synagogue.
Gothic interior of 13th-century Old New Synagogue of Prague.
Historically, synagogues were built in the prevailing architectural style of their time and place. Thus, the synagogue in Kaifeng, China looked very like Chinese temples of that region and era, with its outer wall and open garden in which several buildings were arranged.
Synagogue of the Kaifeng Jewish community in China.
The styles of the earliest synagogues resembled the temples of other sects of the eastern Roman Empire. The synagogues of Morocco are embellished with the colored tilework characteristic of Moroccan architecture. The surviving medieval synagogues in Budapest, Prague and the German lands are typical Gothic structures.
For much of history, the constraints of anti-semitism and the laws of host countries restricting the building of synagogues visible from the street, or forbidding their construction altogether, meant that synagogues were often built within existing buildings, or opening form interior courtyards. In both Europe and in the Muslim world, old synagogues with elaborate interior architecture can be found hidden within nondescript buildings.
Where the building of synagogues was permitted, they were built in the prevailing architectural style of the time and place. Many European cities had elaborate Renaissance synagogues, of which a few survive. In Italy there were many synagogues in the style of the Italian Renaissance (see Leghorn; Padua; and Venice). Those in Padua and Venice possess interiors of great beauty, and are excellent examples of Renaissance work. With the coming of the Baroque era, Baroque synagogues appeared across Europe.
The emancipation of Jews in European countries and of Jews in Muslim countries colonized by European countries gave Jews the right to build large, elaborate synagogues visible from the public street. Synagogue architecture blossomed. Large Jewish communities wished to show not only their wealth but also their newly acquired status as citizens by constructing magnificent synagogues. Handsome nineteenth synagogues form the period of Jewish imagination stand in virtually every country where there were Jewish communities. Most were built in revival styles then in fashion. Thus there were Neoclassical, Neo-Byzantine, Romanesque Revival Moorish Revival, Gothic Revival, and Greek Revival. There are Egyptian Revival synagogues and even one Mayan Revival synagogue. In the nineteenth and early twentieth century heyday of historicist architecture, however, most historicist synagogues, even the most magnificent ones, did not attempt a pure style, or even any particular style, and are best described as eclectic.
Chabad Lubavitch has made a practice of designing some of its Chabad Houses and centers as replicas of or homages to the architecture of 770 Eastern Parkway.
Central Europe: Polish?Lithuanian Commonwealth.
Wolpa Synagogue, 1920, Poland
The great exceptions to the rule that synagogues are built in the prevailing style of their time and place are the wooden synagogues of the former Polish?Lithuanian Commonwealth. They were a unique Jewish artistic and architectural form. Characteristic features include the independence of the pitched roof from the design of the interior domed ceiling. Elaborately carved, painted, domed, balconied and vaulted interiors. The architectural interest of the exterior lay in the large scale of the buildings, the multiple, horizontal lines of the tiered roofs, and the carved corbels that supported them. Wooden synagogues featured a single, large hall. In contrast to contemporary churches, there was no apse. Moreover, while contemporary churches featured imposing vestibules, the entry porches of the wooden synagogues was a low annex, usually with a simple lean-to roof. In these synagogues, the emphasis was on constructing a single, large, high-domed worship space.
According to art historian Stephen S. Kayser, the wooden synagogues of Poland with their painted and carved interiors were "a truly original and organic manifestation of artistic expression?the only real Jewish folk art in history."
According to Louis Lozowick, writing in 1947, the wooden synagogues were unique because, unlike all previous synagogues, they were not built in the architectural style of their region and era, but in a newly evolved and uniquely Jewish style, making them "a truly original folk expression," whose "originality does not lie alone in the exterior architecture, it lies equally in the beautiful and intricate wood carving of the interior."
Moreover, while in many parts of the world Jews were proscribed from entering the building trades and even from practicing the decorative arts of painting and woodcarving, the wooden synagogues were actually built by Jewish craftsmen.
Art historian Ori Z. Soltes points out that the wooden synagogues, unusual for that period in being large, identifiably Jewish buildings not hidden in courtyards or behind walls, were built not only during a Jewish "intellectual golden age" but in a time and place where "the local Jewish population was equal to or even greater than the Christian population.
Egyptian Revival.
Egyptian door, Hobart Synagogue
Egyptian Revival style synagogues were popular in the early nineteenth century. Rachel Wischnitzer argues that they were part of the fashion for Egyptian style inspired by Napoleon's invasion of Europe. According to Carol Herselle Krinsky, they were meant as imitations of the Temple of Solomon and intended by architects and governments to insult Jews by portraying Judaism as a primitive faith. According to Diana Muir Appelbaum, they were expressions of Jewish identity intended to advertise Jewish origins in ancient Israel.
Moorish influence.
Interior of Santa María la Blanca
In medieval Spain (both Al-Andalus and the Christian kingdoms), a host of synagogues were built, and it was usual to commission them from Moorish and later Mudéjar architects. Very few of these medieval synagogues, built with Moorish techniques and style, are conserved. The two best known Spanish synagogues are in Toledo, one known as El Tránsito, the other as Santa María la Blanca, and both, undoubtedly very beautiful, are now preserved as national monuments. The former is a small building containing very rich decorations; the latter is especially noteworthy. It is based upon Almohad style and contains long rows of octagonal columns with curiously carved capitals, from which spring Moorish arches supporting the roof.
Moorish Revival Great Synagogue of Plze?. Another significative Mudéjar synagogue is the one at Córdoba built in 1315. As in El Tránsito, the vegetal and geometrical stucco decorations are purely Moorish, but unlike the former, the epigraphic texts are in Hebrew.
After the expulsion from Spain there was a general feeling among wealthy Sephardim that Moorish architecture was appropriate in synagogues. By the mid-19th century, the style was adopted by the Ashkenazim of Central and Eastern Europe, who associated Moorish and Mudéjar architectural forms with the golden age of Jewry in Al-Andalus. As a consequence, Moorish Revival spread around the globe as a preferred style of synagogue architecture, although Moorish architecture is by no means Jewish, either in fact or in feeling. The Alhambra has furnished inspiration for innumerable synagogues, but seldom have its graceful proportions or its delicate modeling and elaborate ornamentation been successfully copied.
Modern synagogue architecture
In the modern period, synagogues have continued to be built in every popular architectural style, including Art Nouveau, Art Deco, International style, and all contemporary styles. In the post-World War II period "a period of post-war modernism," came to the fore, "characterized by assertive architectural gestures that had the strength and integrity to stand alone, without applied artwork or Jewish iconography." A notable work of Art Nouveau, pre-World War I Hungarian synagogue architecture is Budapest's Kazinczy Street Synagogue.
The most common general plan for the interior of the synagogue is an Ark at the eastern end opposite the entrance, and with an almemar or pulpit. In older or Orthodox synagogues with separate seating, there may be benches for the men on either side, and a women's gallery reached by staircases from the outer vestibule. Variations of this simple plan abound: the vestibule became larger, and the staircases to the women's gallery were separated from the vestibule and given more importance. As the buildings became larger, rows of columns were required to support the roof, but in every case the basilican form was retained. The Ark, formerly allowed a mere niche in the wall, was developed into the main architectural feature of the interior, and was flanked with columns, covered with a canopy and richly decorated. The almemar in many cases was joined to the platform in front of the Ark, and elaborate arrangements of steps were provided.
The Ark.
The Torah Ark (usually called Aron Hakodesh or Hekhál) is the most important feature of the interior, and is generally dignified by proper decoration and raised upon a suitable platform, reached by at least three steps, but often by more. It is usually crowned by the Ten Commandments. The position of the pulpit varies; it may be placed on either side of the Ark, and is occasionally found in the center of the steps.
Other interior arrangements.
The modern synagogue, besides containing the minister's study, trustees' rooms, choir-rooms, and organ-loft, devotes much space to school purposes; generally the entire lower floor is used for class-rooms. The interior treatment of the synagogue allows great latitude in design.
For the thirty-three synagogues of India, American architect and professor of architecture Jay A. Waronker has learned that these buildings tend to follow the Sephardic traditions of the tevah (or bimah, the raised platform where the service is led and Torah read) being freestanding and roughly in the middle of the sanctuary and the ark (called the hekhal by Sephardim and the aron ha-kodesh by Ashkenazim) engaged along the wall that is closest to Jerusalem. The hekhals are essentially cabinets or armoires storing the sefer Torahs. Seating, in the form of long wooden benches, is grouped around and facing the tevah. Men sit together on the main level of the sanctuary while women sit in a dedicated zone on the same level in the smaller synagogues or upstairs in a women's gallery.
Interesting architectural and planning exceptions to this common Sephardic formula are the Cochin synagogues in Kerala of far southwestern India. Here, on the gallery level and adjacent to the space provided for women and overlooking the sanctuary below, is a second tevah. This tevah was used for holidays and unique occasions. It is therefore interesting, on more special events, the woman are closest to the point where the religious service is being led.
In Baghdadi synagogues of India, the hekhals appear to be standard-sized cabinets from the outside (the side facing......
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68.Ants.
Ants are eusocial insects of the family Formicidae /f?r?m?s?di?/ and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from wasp-like ancestors in the mid-Cretaceous period between 110 and 130 million years ago and diversified after the rise of flowering plants. More than 12,500 of an estimated total of 22,000 species have been classified. They are easily identified by their elbowed antennae and the distinctive node-like structure that forms their slender waists.
Ants form colonies that range in size from a few dozen predatory individuals living in small natural cavities to highly organised colonies that may occupy large territories and consist of millions of individuals. Larger colonies consist mostly of sterile, wingless females forming castes of "workers", "soldiers", or other specialised groups. Nearly all ant colonies also have some fertile males called "drones" and one or more fertile females called "queens". The colonies are described as superorganisms because the ants appear to operate as a unified entity, collectively working together to support the colony.
Ants have colonised almost every landmass on Earth. The only places lacking indigenous ants are Antarctica and a few remote or inhospitable islands. Ants thrive in most ecosystems and may form 15?25% of the terrestrial animal biomass. Their success in so many environments has been attributed to their social organisation and their ability to modify habitats, tap resources, and defend themselves. Their long co-evolution with other species has led to mimetic, commensal, parasitic, and mutualistic relationships.
Ant societies have division of labour, communication between individuals, and an ability to solve complex problems. These parallels with human societies have long been an inspiration and subject of study. Many human cultures make use of ants in cuisine, medication, and rituals. Some species are valued in their role as biological pest control agents. Their ability to exploit resources may bring ants into conflict with humans, however, as they can damage crops and invade buildings. Some species, such as the red imported fire ant (Solenopsis invicta), are regarded as invasive species, establishing themselves in areas where they have been introduced accidentally.
The word "ant" is derived from ante, emete of Middle English which are derived from ?mette of Old English, and is related to the dialectal Dutch emt and the Old High German ?meiza, hence the modern German Ameise. All of these words come from West Germanic *?maitij?, and the original meaning of the word was "the biter" (from Proto-Germanic *ai-, "off, away" + *mait- "cut"). The family name Formicidae is derived from the Latin form?ca ("ant") from which the words in other Romance languages, such as the Portuguese formiga, Italian formica, Spanish hormiga, Romanian furnic?, and French fourmi are derived. It has been hypothesised that a Proto-Indo-European word *morwi- was used, cf. Sanskrit vamrah, Latin form?ca, Greek ?????? mýrm?x, Old Church Slavonic mraviji, Old Irish moirb, Old Norse maurr, Dutch mier.
The family Formicidae belongs to the order Hymenoptera, which also includes sawflies, bees, and wasps. Ants evolved from a lineage within the aculeate wasps, and a 2013 study suggests they are a sister group of the Apoidea. In 1966, E. O. Wilson and his colleagues identified the fossil remains of an ant (Sphecomyrma) that lived in the Cretaceous period. The specimen, trapped in amber dating back to around 92 million years ago, has features found in some wasps, but not found in modern ants. Sphecomyrma possibly was a ground forager, while Haidomyrmex and Haidomyrmodes, related genera in subfamily Sphecomyrminae, are reconstructed as active arboreal predators. After the rise of flowering plants about 100 million years ago they diversified and assumed ecological dominance around 60 million years ago. Some groups, such as the Leptanillinae and Martialinae, are suggested to have diversified from early primitive ants that were likely to have been predators underneath the surface of the soil.
During the Cretaceous period, a few species of primitive ants ranged widely on the Laurasian supercontinent (the Northern Hemisphere). They were scarce in comparison to the populations of other insects, representing only about 1% of the entire insect population. Ants became dominant after adaptive radiation at the beginning of the Paleogene period. By the Oligocene and Miocene, ants had come to represent 20?40% of all insects found in major fossil deposits. Of the species that lived in the Eocene epoch, around one in 10 genera survive to the present. Genera surviving today comprise 56% of the genera in Baltic amber fossils (early Oligocene), and 92% of the genera in Dominican amber fossils (apparently early Miocene).
Termites, although sometimes called 'white ants', are not ants. They belong to the sub-order Isoptera within order Blattodea. Termites are more closely related to cockroaches and mantids. Termites are eusocial, but differ greatly in the genetics of reproduction. The similarity of their social structure to that of ants is attributed to convergent evolution. Velvet ants look like large ants, but are wingless female wasps.
Ants are found on all continents except Antarctica, and only a few large islands, such as Greenland, Iceland, parts of Polynesia and the Hawaiian Islands lack native ant species. Ants occupy a wide range of ecological niches, and are able to exploit a wide range of food resources either as direct or indirect herbivores, predators, and scavengers. Most species are omnivorous generalists, but a few are specialist feeders. Their ecological dominance may be measured by their biomass and estimates in different environments suggest that they contribute 15?20% (on average and nearly 25% in the tropics) of the total terrestrial animal biomass, which exceeds that of the vertebrates.
Ants range in size from 0.75 to 52 millimetres (0.030?2.0 in), the largest species being the fossil Titanomyrma giganteum, the queen of which was 6 centimetres (2.4 in) long with a wingspan of 15 centimetres (5.9 in). Ants vary in colour; most ants are red or black, but a few species are green and some tropical species have a metallic lustre. More than 12,000 species are currently known (with upper estimates of the potential existence of about 22,000), with the greatest diversity in the tropics. Taxonomic studies continue to resolve the classification and systematics of ants. Online databases of ant species, including AntBase and the Hymenoptera Name Server, help to keep track of the known and newly described species. The relative ease with which ants may be sampled and studied in ecosystems has made them useful as indicator species in biodiversity studies.
Ants are distinct in their morphology from other insects in having elbowed antennae, metapleural glands, and a strong constriction of their second abdominal segment into a node-like petiole. The head, mesosoma, and metasoma are the three distinct body segments. The petiole forms a narrow waist between their mesosoma (thorax plus the first abdominal segment, which is fused to it) and gaster (abdomen less the abdominal segments in the petiole). The petiole may be formed by one or two nodes (the second alone, or the second and third abdominal segments).
Like other insects, ants have an exoskeleton, an external covering that provides a protective casing around the body and a point of attachment for muscles, in contrast to the internal skeletons of humans and other vertebrates. Insects do not have lungs; oxygen and other gases such as carbon dioxide pass through their exoskeleton via tiny valves called spiracles. Insects also lack closed blood vessels; instead, they have a long, thin, perforated tube along the top of the body (called the "dorsal aorta") that functions like a heart, and pumps haemolymph toward the head, thus driving the circulation of the internal fluids. The nervous system consists of a ventral nerve cord that runs the length of the body, with several ganglia and branches along the way reaching into the extremities of the appendages.
Head.
An ant's head contains many sensory organs. Like most insects, ants have compound eyes made from numerous tiny lenses attached together. Ant eyes are good for acute movement detection, but do not offer a high resolution image. They also have three small ocelli (simple eyes) on the top of the head that detect light levels and polarization. Compared to vertebrates, most ants have poor-to-mediocre eyesight and a few subterranean species are completely blind. Some ants such as Australia's bulldog ant, however, have excellent vision and are capable of discriminating the distance and size of objects moving nearly a metre away.
Two antennae ("feelers") are attached to the head; these organs detect chemicals, air currents, and vibrations; they also are used to transmit and receive signals through touch. The head has two strong jaws, the mandibles, used to carry food, manipulate objects, construct nests, and for defence. In some species a small pocket (infrabuccal chamber) inside the mouth stores food, so it may be passed to other ants or their larvae.
Legs.
All six legs are attached to the mesosoma ("thorax") and terminate in a hooked claw.
Wings.
Only reproductive ants, queens and males, have wings. Queens shed the wings after the nuptial flight, leaving visible stubs, a distinguishing feature of queens. Wingless queens (ergatoids) and males occur in a few species, however.
Metasoma.
The metasoma (the "abdomen") of the ant houses important internal organs, including those of the reproductive, respiratory (tracheae), and excretory systems. Workers of many species have their egg-laying structures modified into stings that are used for subduing prey and defending their nests.
Polymorphism.
Seven Leafcutter ant workers of various castes (left) and two Queens (right)
In the colonies of a few ant species, there are physical castes?workers in distinct size-classes, called minor, median, and major workers. Often, the larger ants have disproportionately larger heads, and correspondingly stronger mandibles. Such individuals are sometimes called "soldier" ants because their stronger mandibles make them more effective in fighting, although they still are workers and their "duties" typically do not vary greatly from the minor or median workers. In a few species, the median workers are absent, creating a sharp divide between the minors and majors. Weaver ants, for example, have a distinct bimodal size distribution. Some other species show continuous variation in the size of workers. The smallest and largest workers in Pheidologeton diversus show nearly a 500-fold difference in their dry-weights. Workers cannot mate; however, because of the haplodiploid sex-determination system in ants, workers of a number of species can lay unfertilised eggs that become fully fertile, haploid males. The role of workers may change with their age and in some species, such as honeypot ants, young workers are fed until their gasters are distended, and act as living food storage vessels. These food storage workers are called repletes. For instance, these replete workers develop in the North American honeypot ant Myrmecocystus mexicanus. Rissing found that usually the largest workers in the colony develop into repletes, and if repletes are removed from the colony other workers become repletes, demonstrating the flexibility of this particular polymorphism. This polymorphism in morphology and behaviour of workers initially was thought to be determined by environmental factors such as nutrition and hormones that led to different developmental paths; however, genetic differences between worker castes have been noted in Acromyrmex sp. These polymorphisms are caused by relatively small genetic changes; differences in a single gene of Solenopsis invicta can decide whether the colony will have single or multiple queens. The Australian jack jumper ant (Myrmecia pilosula) has only a single pair of chromosomes (with the males having just one chromosome as they are haploid), the lowest number known for any animal, making it an interesting subject for studies in the genetics and developmental biology of social insects.
The life of an ant starts from an egg. If the egg is fertilised, the progeny will be female (diploid); if not, it will be male (haploid). Ants develop by complete metamorphosis with the larva stages passing through a pupal stage before emerging as an adult. The larva is largely immobile and is fed and cared for by workers. Food is given to the larvae by trophallaxis, a process in which an ant regurgitates liquid food held in its crop. This is also how adults share food, stored in the "social stomach". Larvae, especially in the later stages, may also be provided solid food such as trophic eggs, pieces of prey, and seeds brought by workers.
The larvae grow through a series of four or five moults and enter the pupal stage. The pupa has the appendages free and not fused to the body as in a butterfly pupa. The differentiation into queens and workers (which are both female), and different castes of workers (when they exist), is influenced in some species by the nutrition the larvae obtain. Genetic influences and the control of gene expression by the developmental environment are complex and the determination of caste continues to be a subject of research. Larvae and pupae need to be kept at fairly constant temperatures to ensure proper development, and so often, are moved around among the various brood chambers within the colony.
A new worker spends the first few days of its adult life caring for the queen and young. She then graduates to digging and other nest work, and later to defending the nest and foraging. These changes are sometimes fairly sudden, and define what are called temporal castes. An explanation for the sequence is suggested by the high casualties involved in foraging, making it an acceptable risk only for ants who are older and are likely to die soon of natural causes.
Most ant species have a system in which only the queen and breeding females have the ability to mate. Contrary to popular belief, some ant nests have multiple queens, while others may exist without queens. Workers with the ability to reproduce are called "gamergates" and colonies that lack queens are then called gamergate colonies; colonies with queens are said to be queen-right. The winged male ants, called drones, emerge from pupae along with the breeding females (although some species, such as army ants, have wingless queens), and do nothing in life except eat and mate.
Most ants are univoltine, producing a new generation each year. During the species-specific breeding period, new reproductives, females and winged males leave the colony in what is called a nuptial flight. Typically, the males take flight before the females. Males then use visual cues to find a common mating ground, for example, a landmark such as a pine tree to which other males in the area converge. Males secrete a mating pheromone that females follow. Females of some species mate with just one male, but in others they may mate with as many as ten or more different males.
Mated females then seek a suitable place to begin a colony. There, they break off their wings and begin to lay and care for eggs. The females store the sperm they obtain during their nuptial flight to selectively fertilise future eggs. The first workers to hatch are weak and smaller than later workers, but they begin to serve the colony immediately. They enlarge the nest, forage for food, and care for the other eggs. This is how new colonies start in most ant species. Species that have multiple queens may have a queen leaving the nest along with some workers to found a colony at a new site, a process akin to swarming in honeybees.
Ants mating.
A wide range of reproductive strategies have been noted in ant species. Females of many species are known to be capable of reproducing asexually through thelytokous parthenogenesis.
Ant colonies can be long-lived. The queens can live for up to 30 years, and workers live from 1 to 3 years. Males, however, are more transitory, being quite short-lived and surviving for only a few weeks. Ant queens are estimated to live 100 times longer than solitary insects of a similar size.
Ants are active all year long in the tropics, but, in cooler regions, they survive the winter in a state of dormancy or inactivity. The forms of inactivity are varied and some temperate species have larvae going into the inactive state, (diapause), while in others, the adults alone pass the winter in a state of reduced activity.
Communication.
Ants communicate with each other using pheromones, sounds, and touch. The use of pheromones as chemical signals is more developed in ants, such as the red harvester ant, than in other hymenopteran groups. Like other insects, ants perceive smells with their long, thin, and mobile antennae. The paired antennae provide information about the direction and intensity of scents. Since most ants live on the ground, they use the soil surface to leave pheromone trails that may be followed by other ants. In species that forage in groups, a forager that finds food marks a trail on the way back to the colony; this trail is followed by other ants, these ants then reinforce the trail when they head back with food to the colony. When the food source is exhausted, no new trails are marked by returning ants and the scent slowly dissipates. This behaviour helps ants deal with changes in their environment. For instance, when an established path to a food source is blocked by an obstacle, the foragers leave the path to explore new routes. If an ant is successful, it leaves a new trail marking the shortest route on its return. Successful trails are followed by more ants, reinforcing better routes and gradually identifying the best path.
Ants use pheromones for more than just making trails. A crushed ant emits an alarm pheromone that sends nearby ants into an attack frenzy and attracts more ants from farther away. Several ant species even use "propaganda pheromones" to confuse enemy ants and make them fight among themselves. Pheromones are produced by a wide range of structures including Dufour's glands, poison glands and glands on the hindgut, pygidium, rectum, sternum, and hind tibia. Pheromones also are exchanged, mixed with food, and passed by trophallaxis, transferring information within the colony. This allows other ants to detect what task group (e.g., foraging or nest maintenance) other colony members belong to. In ant species with queen castes, when the dominant queen stops producing a specific pheromone, workers begin to raise new queens in the colony.
Some ants produce sounds by stridulation, using the gaster segments and their mandibles. Sounds may be used to communicate with colony members or with other species.
Defence.
A Plectroctena sp. attacks another of its kind to protect its territory
See also Insect defences
Ants attack and defend themselves by biting and, in many species, by stinging, often injecting or spraying chemicals, such as formic acid in the case of formicine ants, alkaloids and piperidines in fire ants, and a variety of protein components in other ants. Bullet ants (Paraponera), located in Central and South America, are considered to have the most painful sting of any insect, although it is usually not fatal to humans. This sting is given the highest rating on the Schmidt Sting Pain Index.
The sting of jack jumper ants can be fatal, and an antivenom has been developed for it.
Fire ants, Solenopsis spp., are unique in having a poison sac containing piperidine alkaloids. Their stings are painful and can be dangerous to hypersensitive people.
A weaver ant in fighting position, mandibles wide open
Trap-jaw ants of the genus Odontomachus are equipped with mandibles called trap-jaws, which snap shut faster than any other predatory appendages within the animal kingdom. One study of Odontomachus bauri recorded peak speeds of between 126 and 230 km/h (78 ? 143 mph), with the jaws closing within 130 microseconds on average. The ants were also observed to use their jaws as a catapult to eject intruders or fling themselves backward to escape a threat. Before striking, the ant opens its mandibles extremely widely and locks them in this position by an internal mechanism. Energy is stored in a thick band of muscle and explosively released when triggered by the stimulation of sensory organs resembling hairs on the inside of the mandibles. The mandibles also permit slow and fine movements for other tasks. Trap-jaws also are seen in the following genera: Anochetus, Orectognathus, and Strumigenys, plus some members of the Dacetini tribe, which are viewed as examples of convergent evolution.
A Malaysian species of ant in the Camponotus cylindricus group has enlarged mandibular glands that extend into their gaster. When disturbed, workers rupture the membrane of the gaster, causing a burst of secretions containing acetophenones and other chemicals that immobilise small insect attackers. The worker subsequently dies.
Suicidal defences by workers are also noted in a Brazilian ant, Forelius pusillus, where a small group of ants leaves the security of the nest after sealing the entrance from the outside each evening.
Ant mound holes prevent water from entering the nest during rain
In addition to defence against predators, ants need to protect their colonies from pathogens. Some worker ants maintain the hygiene of the colony and their activities include undertaking or necrophory, the disposal of dead nest-mates. Oleic acid has been identified as the compound released from dead ants that triggers necrophoric behaviour in Atta mexicana while workers of Linepithema humile react to the absence of characteristic chemicals (dolichodial and iridomyrmecin) present on the cuticle of their living nestmates to trigger similar behaviour.
Nests may be protected from physical threats such as flooding and overheating by elaborate nest architecture. Workers of Cataulacus muticus, an arboreal species that lives in plant hollows, respond to flooding by drinking water inside the nest, and excreting it outside. Camponotus anderseni, which nests in the cavities of wood in mangrove habitats, deals with submergence under water by switching to anaerobic respiration.
Learning.
Many animals can learn behaviours by imitation, but ants may be the only group apart from mammals where interactive teaching has been observed. A knowledgeable forager of Temnothorax albipennis will lead a naive nest-mate to newly discovered food by the process of tandem running. The follower obtains knowledge through its leading tutor. The leader is acutely sensitive to the progress of the follower and slows down when the follower lags and speeds up when the follower gets too close.
Controlled experiments with colonies of Cerapachys biroi suggest that an individual may choose nest roles based on her previous experience. An entire generation of identical workers was divided into two groups whose outcome in food foraging was controlled. One group was continually rewarded with prey, while it was made certain that the other failed. As a result, members of the successful group intensified their foraging attempts while the unsuccessful group ventured out fewer and fewer times. A month later, the successful foragers continued in their role while the others had moved to specialise in brood care.
Nest construction.
Complex nests are built by many ant species, but other species are nomadic and do not build permanent structures. Ants may form subterranean nests or build them on trees. These nests may be found in the ground, under stones or logs, inside logs, hollow stems, or even acorns. The materials used for construction include soil and plant matter, and ants carefully select their nest sites; Temnothorax albipennis will avoid sites with dead ants, as these may indicate the presence of pests or disease. They are quick to abandon established nests at the first sign of threats.
The army ants of South America, such as the Eciton burchellii species, and the driver ants of Africa do not build permanent nests, but instead, alternate between nomadism and stages where the workers form a temporary nest (bivouac) from their own bodies, by holding each other together.
Weaver ant (Oecophylla spp.) workers build nests in trees by attaching leaves together, first pulling them together with bridges of workers and then inducing their larvae to produce silk as they are moved along the leaf edges. Similar forms of nest construction are seen in some species of Polyrhachis.
Formica polyctena, among other ant species, constructs nests that maintain a relatively constant interior temperature that aids in the development of larvae. The ants maintain the nest temperature by choosing the location, nest materials, controlling ventilation and maintaining the heat from solar radiation, worker activity and metabolism, and in some moist nests, microbial activity in the nest materials.
Some ant species, such as those that use natural cavities, can be opportunistic and make use of the controlled micro-climate provided inside human dwellings and other artificial structures to house their colonies and nest structures.
Cultivation of food.
Myrmecocystus, honeypot ants, store food to prevent colony famine
Most ants are generalist predators, scavengers, and indirect herbivores, but a few have evolved specialised ways of obtaining nutrition. It is believed that many ant species that engage in indirect herbivory rely on specialized symbiosis with their gut microbes to upgrade the nutritional value of the food they collect and allow them to survive in nitrogen poor regions, such as rainforrest canopies. Leafcutter ants (Atta and Acromyrmex) feed exclusively on a fungus that grows only within their colonies. They continually collect leaves which are taken to the colony, cut into tiny pieces and placed in fungal gardens. Workers specialise in related tasks according to their sizes. The largest ants cut stalks, smaller workers chew the leaves and the smallest tend the fungus. Leafcutter ants are sensitive enough to recognise the reaction of the fungus to different plant material, apparently detecting chemical signals from the fungus. If a particular type of leaf is found to be toxic to the fungus, the colony will no longer collect it. The ants feed on structures produced by the fungi called gongylidia. Symbiotic bacteria on the exterior surface of the ants produce antibiotics that kill bacteria introduced into the nest that may harm the fungi.
Navigation.
Foraging ants travel distances of up to 200 metres (700 ft) from their nest and scent trails allow them to find their way back even in the dark. In hot and arid regions, day-foraging ants face death by desiccation, so the ability to find the shortest route back to the nest reduces that risk. Diurnal desert ants of the genus Cataglyphis such as the Sahara desert ant navigate by keeping track of direction as well as distance travelled. Distances travelled are measured using an internal pedometer that keeps count of the steps taken and also by evaluating the movement of objects in their visual field (optical flow). Directions are measured using the position of the sun. They integrate this information to find the shortest route back to their nest. Like all ants, they can also make use of visual landmarks when available as well as olfactory and tactile cues to navigate. Some species of ant are able to use the Earth's magnetic field for navigation. The compound eyes of ants have specialised cells that detect polarised light from the Sun, which is used to determine direction. These polarization detectors are sensitive in the ultraviolet region of the light spectrum. In some army ant species, a group of foragers who become separated from the main column sometimes may turn back on themselves and form a circular ant mill. The workers may then run around continuously until they die of exhaustion.
Locomotion.
The female worker ants do not have wings and reproductive females lose their wings after their mating flights in order to begin their colonies. Therefore, unlike their wasp ancestors, most ants travel by walking. Some species are capable of leaping. For example, Jerdon's jumping ant (Harpegnathos saltator) is able to jump by synchronising the action of its mid and hind pairs of legs. There are several species of gliding ant including Cephalotes atratus; this may be a common trait among most arboreal ants. Ants with this ability are able to control the direction of their descent while falling.
Other species of ants can form chains to bridge gaps over water, underground, or through spaces in vegetation. Some species also form floating rafts that help them survive floods. These rafts may also have a role in allowing ants to colonise islands. Polyrhachis sokolova, a species of ant found in Australian mangrove swamps, can swim and live in underwater nests. Since they lack gills, they go to trapped pockets of air in the submerged nests to breathe.
Cooperation and competition.
Meat-eater ants feeding on a cicada, social ants cooperate and collectively gather food
Not all ants have the same kind of societies. The Australian bulldog ants are among the biggest and most basal of ants. Like virtually all ants, they are eusocial, but their social behaviour is poorly developed compared to other species. Each individual hunts alone, using her large eyes instead of chemical senses to find prey.
Some species (such as Tetramorium caespitum) attack and take over neighbouring ant colonies. Others are less expansionist, but just as aggressive; they invade colonies to steal eggs or larvae, which they either eat or raise as workers or slaves. Extreme specialists among these slave-raiding ants, such as the Amazon ants, are incapable of feeding themselves and need captured workers to survive. Captured workers of the enslaved species Temnothorax have evolved a counter strategy, destroying just the female pupae of the slave-making Protomognathus americanus, but sparing the males (who don't take part in slave-raiding as adults).
A worker Harpegnathos saltator (a jumping ant) engaged in battle with a rival colony's queen
Ants identify kin and nestmates through their scent, which comes from hydrocarbon-laced secretions that coat their exoskeletons. If an ant is separated from its original colony, it will eventually lose the colony scent. Any ant that enters a colony without a matching scent will be attacked. Also, the reason why two separate colonies of ants will attack each other even if they are of the same species is because the genes responsible for pheromone production are different between them. The Argentine ant, however, does not have this characteristic, due to lack of genetic diversity, and has become a global pest because of it.
Parasitic ant species enter the colonies of host ants and establish themselves as social parasites; species such as Strumigenys xenos are entirely parasitic and do not have workers, but instead, rely on the food gathered by their Strumigenys perplexa hosts. This form of parasitism is seen across many ant genera, but the parasitic ant is usually a species that is closely related to its host. A variety of methods are employed to enter the nest of the host ant. A parasitic queen may enter the host nest before the first brood has hatched, establishing herself prior to development of a colony scent. Other species use pheromones to confuse the host ants or to trick them into carrying the parasitic queen into the nest. Some simply fight their way into the nest.
A conflict between the sexes of a species is seen in some species of ants with these reproducers apparently competing to produce offspring that are as closely related to them as possible. The most extreme form involves the production of clonal offspring. An extreme of sexual conflict is seen in Wasmannia auropunctata, where the queens produce diploid daughters by thelytokous parthenogenesis and males produce clones by a process whereby a diploid egg loses its maternal contribution to produce haploid males who are clones of the father.
Relationships with other organisms.
The spider Myrmarachne plataleoides (female shown) mimics weaver ants to avoid predators.
Ants form symbiotic associations with a range of species, including other ant species, other insects, plants, and fungi. They also are preyed on by many animals and even certain fungi. Some arthropod species spend part of their lives within ant nests, either preying on ants, their larvae, and eggs, consuming the food stores of the ants, or avoiding predators. These inquilines may bear a close resemblance to ants. The nature of this ant mimicry (myrmecomorphy) varies, with some cases involving Batesian mimicry, where the mimic reduces the risk of predation. Others show Wasmannian mimicry, a form of mimicry seen only in inquilines.
Aphids and other hemipteran insects secrete a sweet liquid called honeydew, when they feed on plant sap. The sugars in honeydew are a high-energy food source, which many ant species collect. In some cases, the aphids secrete the honeydew in response to ants tapping them with their antennae. The ants in turn keep predators away from the aphids and will move them from one feeding location to another. When migrating to a new area, many colonies will take the aphids with them, to ensure a continued supply of honeydew. Ants also tend mealybugs to harvest their honeydew. Mealybugs may become a serious pest of pineapples if ants are present to protect mealybugs from their natural enemies.
Myrmecophilous (ant-loving) caterpillars of the butterfly family Lycaenidae (e.g., blues, coppers, or hairstreaks) are herded by the ants, led to feeding areas in the daytime, and brought inside the ants' nest at night. The caterpillars have a gland which secretes honeydew when the ants massage them. Some caterpillars produce vibrations and sounds that are perceived by the ants. Other caterpillars have evolved from ant-loving to ant-eating: these myrmecophagous caterpillars secrete a pheromone that makes the ants act as if the caterpillar is one of their own larvae. The caterpillar is then taken into the ant nest where it feeds on the ant larvae. Fungus-growing ants that make up the tribe Attini, including leafcutter ants, cultivate certain species of fungus in the Leucoagaricus or Leucocoprinus genera of the Agaricaceae family. In this ant-fungus mutualism, both species depend on each other for survival. The ant Allomerus decemarticulatus has evolved a three-way association with the host plant, Hirtella physophora (Chrysobalanaceae), and a sticky fungus which is used to trap their insect prey.
Ants may obtain nectar from flowers such as the dandelion but are only rarely known to pollinate flowers.
Lemon ants make devil's gardens by killing surrounding plants with their stings and leaving a pure patch of lemon ant trees, (Duroia hirsuta). This modification of the forest provides the ants with more nesting sites inside the stems of the Duroia trees. Although some ants obtain nectar from flowers, pollination by ants is somewhat rare. Some plants have special nectar exuding structures, extrafloral nectaries that provide food for ants, which in turn protect the plant from more damaging herbivorous insects. Species such as the bullhorn acacia (Acacia cornigera) in Central America have hollow thorns that house colonies of stinging ants (Pseudomyrmex ferruginea) who defend the tree against insects, browsing mammals, and epiphytic vines. Isotopic labelling studies suggest that plants also obtain nitrogen from the ants. In return, the ants obtain food from protein- and lipid-rich Beltian bodies. Another example of this type of ectosymbiosis comes from the Macaranga tree, which has stems adapted to house colonies of Crematogaster ants.
Many tropical tree species have seeds that are dispersed by ants. Seed dispersal by ants or myrmecochory is widespread and new estimates suggest that nearly 9% of all plant species may have such ant associations. Some plants in fire-prone grassland systems are particularly dependent on ants for their survival and dispersal as the seeds are transported to safety below the ground. Many ant-dispersed seeds have special external structures, elaiosomes, that are sought after by ants as food.
A convergence, possibly a form of mimicry, is seen in the eggs of stick insects. They have an edible elaiosome-like structure and are taken into the ant nest where the young hatch.
Most ants are predatory and some prey on and obtain food from other social insects including other ants. Some species specialise in preying on termites (Megaponera and Termitopone) while a few Cerapachyinae prey on other ants. Some termites, including Nasutitermes corniger, form associations with certain ant species to keep away predatory ant species. The tropical wasp Mischocyttarus drewseni coats the pedicel of its nest with an ant-repellent chemical. It is suggested that many tropical wasps may build their nests in trees and cover them to protect themselves from ants. Other wasps such as A. multipicta defend against ants by blasting them off the nest with bursts of wing buzzing. Stingless bees (Trigona and Melipona) use chemical defences against ants. Certain species of ants have the power to drive certain wasps, such as Polybia occidentalis to extinction if they attack more than once and the wasps cannot keep up with rebuilding their nest.
Flies in the Old World genus Bengalia (Calliphoridae) prey on ants and are kleptoparasites, snatching prey or brood from the mandibles of adult ants. Wingless and legless females of the Malaysian phorid fly (Vestigipoda myrmolarvoidea) live in the nests of ants of the genus Aenictus and are cared for by the ants.
Fungi in the genera Cordyceps and Ophiocordyceps infect ants. Ants react to their infection by climbing up plants and sinking their mandibles into plant tissue. The fungus kills the ants, grows on their remains, and produces a fruiting body. It appears that the fungus alters the behaviour of the ant to help disperse its spores in a microhabitat that best suits the fungus. Strepsipteran parasites also manipulate their ant host to climb grass stems, to help the parasite find mates.
A nematode (Myrmeconema neotropicum) that infects canopy ants (Cephalotes atratus) causes the black-coloured gasters of workers to turn red. The parasite also alters the behaviour of the ant, causing them to carry their gasters high. The conspicuous red gasters are mistaken by birds for ripe fruits, such as Hyeronima alchorneoides, and eaten. The droppings of the bird are collected by other ants and fed to their young, leading to further spread of the nematode.
South American poison dart frogs in the genus Dendrobates feed mainly on ants, and the toxins in their skin may come from the ants.
Army ants forage in a wide roving column, attacking any animals in that path that are unable to escape. In Central and South America, Eciton burchellii is the swarming ant most commonly attended by "ant-following" birds such as antbirds and woodcreepers. This behaviour was once considered mutualistic, but later studies found the birds to be parasitic. Although direct kleptoparasitism (birds stealing food from the ants' grasp) is rare, the birds eat many prey insects that the ants would otherwise eat and thus decrease their foraging success. Birds indulge in a peculiar behaviour called anting that, as yet, is not fully understood. Here birds rest on ant nests, or pick and drop ants onto their wings and feathers; this may be a means to remove ectoparasites from the birds.
Anteaters, aardvarks, pangolins, echidnas and numbats have special adaptations for living on a diet of ants. These adaptations include long, sticky tongues to capture ants and strong claws to break into ant nests. Brown bears (Ursus arctos) have been found to feed on ants. About 12%, 16%, and 4% of their faecal volume in spring, summer, and autumn, respectively, is composed of ants.
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